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Assimineidae is a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of minute
snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class G ...
s, also known as palmleaf snails, with an operculum, gastropod
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is es ...
s or
micromollusk A micromollusk is a shelled mollusk which is extremely small, even at full adult size. The word is usually, but not exclusively, applied to marine mollusks, although in addition, numerous species of land snails and freshwater mollusks also ...
s in the superfamily Rissoidae. Many of these very small snails live in intermediate habitats, being amphibious between saltwater and land; others live in freshwater.


Distribution

The distribution of the Assimineidae is worldwide."Family summary for Assimineidae"
AnimalBase AnimalBase is a project brought to life in 2004 and is maintained by the University of Göttingen, Germany. The goal of the AnimalBase project is to digitize early zoological literature, provide copyright-free open access to zoological works, and p ...
, last change 15-08-2006, accessed 4 August 2010.
The oldest fossils are known from the Cenomanian aged
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. ...
.


Ecology

Various species in this family occur in
estuarine An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environmen ...
habitats, in
salt marsh A salt marsh or saltmarsh, also known as a coastal salt marsh or a tidal marsh, is a coastal ecosystem in the upper coastal intertidal zone between land and open saltwater or brackish water that is regularly flooded by the tides. It is domin ...
es and in freshwater. Some are terrestrial or
amphibious Amphibious means able to use either land or water. In particular it may refer to: Animals * Amphibian, a vertebrate animal of the class Amphibia (many of which live on land and breed in water) * Amphibious caterpillar * Amphibious fish, a fish ...
.


Description

The
shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses ** Thin-shell structure Science Biology * Seashell, a hard o ...
is small to medium large, more or less egg-conelike shaped. The apertural margin is simple. The operculum is in most cases horny. Species in this family are characterized by rudimentary cephalic
tentacles In zoology, a tentacle is a flexible, mobile, and elongated organ present in some species of animals, most of them invertebrates. In animal anatomy, tentacles usually occur in one or more pairs. Anatomically, the tentacles of animals work mai ...
, a trunklike
snout A snout is the protruding portion of an animal's face, consisting of its nose, mouth, and jaw. In many animals, the structure is called a muzzle, rostrum, or proboscis. The wet furless surface around the nostrils of the nose of many mammals is ...
, a foot with a groove and rudimentary to absent ctenidium (a comb-like respiratory apparatus).


Taxonomy

The following three
subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
were recognized in the taxonomy of Bouchet & Rocroi (2005): * Subfamily Assimineinae H. Adams & A. Adams, 1856 - synonym: Synceratidae Bartsch, 1920 * Subfamily Ekadantinae Thiele, 1929 - synonyms: Paludinellidae Kobelt, 1878 (n.a.); Cyclotropidae Iredale, 1941 * Subfamily Omphalotropidinae Thiele, 1927 - Realiinae L. Pfeiffer, 1853 (inv.); Adelomorphinae Kobelt, 1906 (inv.); Garrettiinae Kobelt, 1906; Pseudocyclotini Thiele, 1929; Thaanumellinae Clench, 1946; Tutuilanidae Hubendick, 1952


Genera

Genera in the family Assimineidae include:ZipCodeZoo
Genus † '' Laternoides'' W. Yu & Y.-H. Xi, 1977 subfamily Assimineinae * '' Angustassiminea'' Habe, 1943 * '' Assiminea'' Fleming, 1828 - type genus of Assimineidae, synonym: ''Assiminia'' * '' Austropilula'' Thiele, 1927 * '' Aviassiminea'' Fukuda & Ponder, 2003 * '' Conassiminea'' Fukuda & Ponder, 2006 * '' Cryptassiminea'' Fukuda & Ponder, 2005 * '' Euassiminea'' Heude, 1882 * '' Macrassiminea'' Thiele, 1927 * '' Ovassiminea'' Thiele, 1927 * '' Pseudomphala'' Heude, 1882 * '' Sculptassiminea'' Thiele, 1927 subfamily Ekadantinae * '' Acmella'' Blanford, 1869 * '' Cavernacmella'' Habe, 1942 * '' Cyclotropis'' Tapparone Canefri, 1883 * '' Ekadanta'' Rao, 1928 * '' Metassiminea'' Thiele, 1927 * ''
Optediceros ''Optediceros'' is a genus of minute operculate snails, marine gastropod mollusks or micromollusks in the subfamily Ekadantinae of the family Assimineidae.MolluscaBase (2019). MolluscaBase. Optediceros Leith, 1853. Accessed through: World Regist ...
'' Leith, 1853 * ''
Paludinella ''Paludinella'' is a genus of minute salt marsh snails with an Operculum (gastropod), operculum, Aquatic animal, aquatic gastropod mollusks or micromollusks, in the family Assimineidae.Bouchet, P.; Gofas, S. (2012). Paludinella Pfeiffer, 1841. Ac ...
'' Pfeiffer, 1841 * '' Rugapedia'' Fukuda & Ponder, 2004 * '' Solenomphala'' Heude, 1882 * '' Taiwanassiminea'' Kuroda & Habe, 1950 subfamily Omphalotropidinae Thiele, 1927 (synonym: Garrettiinae Kobelt, 1906) * '' Allepithema'' Tomlin, 1931 * '' Anaglyphula'' B. Rensch, 1932 * ''
Atropis Assimineidae is a family of minute snails, also known as palmleaf snails, with an operculum, gastropod mollusks or micromollusks in the superfamily Rissoidae. Many of these very small snails live in intermediate habitats, being amphibious betwee ...
'' Pease, 1871 * ''
Austroassiminea ''Austroassiminea'' is a genus of minute, salt marsh snails with an operculum, aquatic gastropod mollusks, or micromollusks, in the family Assimineidae. Species Species in the genus ''Austroassiminea'': * ''Austroassiminea letha ''Austro ...
'' Solem, Girardi, Slack-Smith & Kendrick, 1982 * '' Balambania'' Crosse, 1891 * '' Chalicopoma'' Möllendorff, 1894 * ''
Conacmella ''Conacmella'' is a genus of minute, salt marsh snails with an operculum, aquatic gastropod mollusks, or micromollusks, in the family Assimineidae Assimineidae is a family of minute snails, also known as palmleaf snails, with an operculum, ...
'' Thiele, 1925 * '' Crossilla'' Thiele, 1927 * ''
Cyclomorpha ''Cyclaxyra'' is a genus of cucujoid beetles in the family Cyclaxyridae, and the sole extant genus in the family, others being known only from fossils.Gimmel, M.L., Szawaryn K., Cai C. & Leschen R.A.B. (2019) Mesozoic sooty mould beetles as livi ...
'' Pease, 1871 * '' Ditropisena'' Iredale, 1933 * '' Duritropis'' Iredale, 1944 * '' Electrina'' Baird, 1850 * '' Eussoia'' Preston, 1912 * '' Fijianella'' C. M. Cooke & Clench, 1943 * ''
Garrettia ''Garrettia'' is a genus of flowering plant in the family Lamiaceae, first described in 1937. It contains only one known species, ''Garrettia siamensis''. It is native to southern China (Yunnan Province), Java, Thailand Thailand ( ), ...
'' Paetel, 1873 * '' Kubaryia'' Clench, 1948 * '' Leucostele'' Thiele, 1927 * ''
Limborelia ''Limborelia'' is a genus in the subfamily Omphalotropidinae of palmleaf snails that is endemic to Australia's Lord Howe Island in the Tasman Sea The Tasman Sea ( Māori: ''Te Tai-o-Rēhua'', ) is a marginal sea of the South Pacific O ...
'' Iredale, 1944 * '' Nesopoma'' Clench, 1958 * ''
Omphalotropis ''Omphalotropis'' is a genus of minute salt marsh snails with an operculum, terrestrial gastropod mollusks in the subfamily Omphalotropidinae of the family Assimineidae.MolluscaBase eds. (2021). MolluscaBase. Omphalotropis L. Pfeiffer, 1851. ...
'' Pfeiffer, 1851 - type genus of the subfamily Omphalotropidinae * '' Opinorelia'' Iredale, 1944 * '' Paludinellassiminea'' Habe, 1994 * '' Ponapella'' Clench, 1946 * '' Pseudassiminea'' Thiele, 1927 * '' Pseudocyclotus'' Thiele, 1894 * '' Pseudogibbula'' Dautzenberg, 1890 * '' Quadrasiella'' Moellendorff, 1894 * '' Rapanella'' Cooke & Clench, 1943 * '' Rupacilla'' Thiele, 1927 * '' Scalinella'' Pease, 1867 * '' Schuettiella'' Brandt, 1974 * ''
Setaepoma ''Setaepoma'' is a genus of minute salt marsh snails with an operculum, terrestrial gastropod mollusks in the subfamily Omphalotropidinae of the family Assimineidae Assimineidae is a family of minute snails, also known as palmleaf snails, ...
'' Clench, 1955 * '' Spiratropis'' Kobelt & Möllendorff, 1900 * ''
Suterilla ''Suterilla'' is a genus of minute operculate snails, marine gastropod mollusks or micromollusks in the family Assimineidae. Species Species within the genus ''Suterilla'' include: * ''Suterilla climoi'' Fukuda, Ponder & Marshall, 2006 * ...
'' Thiele, 1927 * '' Sychnotropis'' Möllendorff, 1898 * '' Telmosena'' Iredale, 1944 * '' Thaanumella'' Clench, 1946 * '' Turbacmella'' Thiele, 1927 * '' Tutuilana'' Hubendick, 1952 * ''
Wrayanna ''Wrayanna'' is a genus of minute, salt marsh snails with an operculum, aquatic gastropod mollusks, or micromollusks, in the family Assimineidae Assimineidae is a family of minute snails, also known as palmleaf snails, with an operculum, g ...
'' Clench, 1948 ;incertae sedis: * Assimineidae incertae sedis vulgaris Webster, 1905 Rosenberg, G. (2012). Assimineidae incertae sedis vulgaris. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=598622 on 2012-05-31 * ''Assimineidae incertae sedis turrita'' (W. H. Turton, 1932) (taxon inquirendum) * ''Assimineidae incertae sedis turritella'' (W. H. Turton, 1932) (taxon inquirendum) ;Synonyms: * ''Adelomorpha'' Tapparone Canefri, 1886: synonym of ''Pseudocyclotus'' Thiele, 1894 (Invalid: Junior homonym of ''Adelomorpha'' Snellen, 1885 epidoptera * ''Adelostoma'' E. A. Smith, 1885: synonym of ''Pseudocyclotus'' Thiele, 1894 (invalid: junior homonym of ''Adelostoma'' Duponchel, 1827) * ''Assemania'' Dollfus, 1912: synonym of ''Assiminea'' J. Fleming, 1828 (unjustified emendation) * ''Assimania'': synonym of ''Assiminea'' J. Fleming, 1828 (incorrect spelling of genus name) * ''Assiminella'' Monterosato, 1906: synonym of ''Paludinella'' L. Pfeiffer, 1841 (unnecessary substitute name for Paludinella L. Pfeiffer, 1841) * ''Assiminia'' J. Fleming, 1828: synonym of ''Assiminea'' J. Fleming, 1828 (alternative original spelling
sed in the index p. 557 sed ("stream editor") is a Unix utility that parses and transforms text, using a simple, compact programming language. It was developed from 1973 to 1974 by Lee E. McMahon of Bell Labs, and is available today for most operating systems. sed ...
* ''Diadema'' Pease, 1868: synonym of ''Garrettia'' Paetel, 1873 (invalid: junior homonym of ''Diadema'' Gray, 1825) * ''Dramelia'' Iredale, 1941: synonym of ''Acmella'' W. T. Blanford, 1869 (junior synonym) * ''Heteropoma'' Möllendorff, 1894: synonym of ''Allepithema'' Tomlin, 1931 (invalid: junior homonym of ''Heteropoma'' Benson, 1856; ''Allepithema'' is a replacement name) * ''Paludinassiminea'' Matsubayashi & Habe, 1990: synonym of ''Solenomphala'' Heude, 1882 (unavailable name: no type species designated) * Tribe ''Pseudocyclotini'' Thiele, 1929: synonym of ''Omphalotropidinae'' Thiele, 1927 * ''Scalinella'' Pease, 1868: synonym of ''Omphalotropis'' L. Pfeiffer, 1851 * ''Stenotropis'' Kobelt & Möllendorff, 1898: synonym of ''Omphalotropis (Stenotropis)'' Kobelt & Möllendorff, 1898 represented as ''Omphalotropis'' L. Pfeiffer, 1851 * ''Syncera'' Gray, 1821 (nomen nudum)


References

This article incorporates public domain text from the reference.


External links

{{Taxonbar, from=Q3171053 pt:Assiminea