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Assamiidae are a family of harvestmen with more than 400 described species. It is the third most diverse family of the suborder
Laniatores Laniatores is the largest suborder of the arachnid order Opiliones with over 4,200 described species worldwide. The majority of the species are highly dependent on humid environments and usually correlated with tropical and temperate forest habit ...
.


Name

The family name is derived from the
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
n province
Assam Assam (, , ) is a state in Northeast India, northeastern India, south of the eastern Himalayas along the Brahmaputra Valley, Brahmaputra and Barak River valleys. Assam covers an area of . It is the second largest state in Northeast India, nor ...
, where the type species of the type genus ('' Assamia westermanni'') was collected. (2007): Assamiidae Sørensen, 1884. In: Pinto-da-Rocha ''et al.'' 2007: 173ff


Description

These harvestmen range in body length from two to eight millimeters. The length of their legs ranges from four to forty millimeters, though they are usually long. Assamiidae are usually reddish brown to yellow with black mottling and reticulation. Some species have white drawings on the dorsal scutum.


Distribution

Assamiidae only occur in the
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
, and are completely absent from
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
and
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
, with most species found in
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
and southern
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
. Although they are also not found on the
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is bounded by the cont ...
islands, the subfamily Dampetrinae has radiated in
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
,
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
, and somewhat in
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
.


Subtaxa

A list of included species is
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Although Roewer established numerous subfamilies for this group, most of these are unsupported. There are at least five great groups, whose boundaries do not coincide with these subfamilies. :The Dampetrinae are endemic to Australia and New Guinea. :The Assamiinae occur mostly in
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
and
Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
. :The Trionyxellinae have a pseudonychium (a median
tarsal claw This glossary describes the terms used in formal descriptions of spiders; where applicable these terms are used in describing other arachnids. Links within the glossary are shown . Terms A * Abdomen or opisthosoma: One of the two main body par ...
on the third and fourth leg tarsi) and are found in India and
Sri Lanka Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia. It lies in the Indian Ocean, southwest of the Bay of Bengal, separated from the Indian subcontinent, ...
. :The Erecinae live in central Africa. :The small, blind Irumuinae dwell in
caves Caves or caverns are natural voids under the Earth's surface. Caves often form by the weathering of rock and often extend deep underground. Exogene caves are smaller openings that extend a relatively short distance underground (such as rock ...
and
soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...
. For a list of currently described species with the traditional groupings into subfamilies, see the List of Assamiidae species.


Relationships

The Assamidae are possibly the sister group to the
Gonyleptoidea Gonyleptoidea is the most diverse superfamily of the Grassatores. It includes around 2,500 species distributed in the Neotropics. They are characterized by the simplified male genitalia, with the glans free subapical in the truncus. Gonyleptoide ...
. The spiny funnel on the
penis A penis (; : penises or penes) is a sex organ through which male and hermaphrodite animals expel semen during copulation (zoology), copulation, and through which male placental mammals and marsupials also Urination, urinate. The term ''pen ...
is very similar to those of Stygnopsidae from
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
. Mello-Leitão erected the Trionyxellidae for four subfamilies of Assamiidae in 1949, but this was ignored by later authors.


Footnotes


References

* (eds.) (2007): Harvestmen - The Biology of Opiliones. ''Harvard University Press'' {{Taxonbar, from=Q142563 Harvestman families