Asiatheriidae
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''Asiatherium'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
, probably belonging to
Metatheria Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as wel ...
. It lived during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
, and its fossilized remains were discovered in
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
.


Description

Skull of ''Asiatherium'' is in length. This animal was roughly the size of a
mouse A mouse (: mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus'' ...
, and is known from a fairly complete articulated skeleton, preserving a complete skull. Its teeth resembled roughly those of
marsupial Marsupials are a diverse group of mammals belonging to the infraclass Marsupialia. They are natively found in Australasia, Wallacea, and the Americas. One of marsupials' unique features is their reproductive strategy: the young are born in a r ...
s; it possessed three premolars and four molars, and the mandible was slightly curved. Compared to '' Deltatheridium'', ''Asiatherium'' was even more marsupial-like, due to its molars having paraconids lower than the metaconids, and the paired entoconid-hypoconulid cusps on the lower molars. The stylar platform of the upper molars was however narrower and the stylar cusps weaker than in primitive
marsupial Marsupials are a diverse group of mammals belonging to the infraclass Marsupialia. They are natively found in Australasia, Wallacea, and the Americas. One of marsupials' unique features is their reproductive strategy: the young are born in a r ...
s. The upper molars differed from those of marsupials in their expanded praecingula and postcingula.*F. S. Szalay and B. A. Trofimov. 1996. The Mongolian Late Cretaceous Asiatherium, and the early phylogeny and paleobiogeography of Metatheria. Journal of Vertebrate Paleontology 16(3):474-509


Classification

''Asiatherium reshetovi'' was first described in 1994, based on fossils found in the Udan-Sayr locality of the
Djadokhta Formation The Djadochta formation (sometimes transcribed and also known as Djadokhta, Djadokata, or Dzhadokhtskaya) is a highly fossiliferous geological formation in Central Asia, Gobi Desert, dating from the Late Cretaceous period, about 75 million ...
, in
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
. ''Asiatherium'' seems to be related to early marsupials, but probably was nested within a different clade of
Metatherian Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well ...
,
Deltatheroida Deltatheroida is an extinct group of basal metatherians that were distantly related to modern marsupials. The majority of known members of the group lived in the Cretaceous; one species, '' Gurbanodelta kara'', is known from the late Paleocene ( ...
, nested alongside Deltatheridium. The skeleton of ''Asiatherium'' looked like that of a basal
Theria Theria ( or ; ) is a scientific classification, subclass of mammals amongst the Theriiformes. Theria includes the eutherians (including the Placentalia, placental mammals) and the metatherians (including the marsupials) but excludes the egg-lay ...
n; epipubic bones, typical of marsupials, were present.


References and Bibliography

*B. A. Trofimov and F. S. Szalay. 1994. New Cretaceous marsupial from Mongolia and the early radiation of Metatheria. Proceedings of the National Academy of Sciences 91:12569-12573 *T. E. Williamson, S. L. Brusatte, T. D. Carr, A. Weil, and B. R. Standhardt. 2012. The phylogeny and evolution of Cretaceous–Palaeogene metatherians: cladistic analysis and description of new early Palaeocene specimens from the Nacimiento Formation, New Mexico. Journal of Systematic Palaeontology 10(4):625-651 {{Taxonbar, from=Q30895124 Cretaceous mammals Cretaceous mammals of Asia Monotypic prehistoric mammal genera Fossil taxa described in 1994 Prehistoric metatherians