Argyrosaurus Superbus
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''Argyrosaurus'' ( ) is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
titanosaurian Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of Sauropoda, sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with tax ...
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
that lived about 70
million years ago Million years ago, abbreviated as Mya, Myr (megayear) or Ma (megaannum), is a unit of time equal to (i.e. years), or approximately 31.6 teraseconds. Usage Myr is in common use in fields such as Earth science and cosmology. Myr is also used w ...
, during the Late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
Period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...
of what is now
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
.


Discovery and naming

The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, ''Argyrosaurus superbus'', was formally described by
Richard Lydekker Richard Lydekker (; 25 July 1849 – 16 April 1915) was a British naturalist, geologist and writer of numerous books on natural history. He was known for his contributions to zoology, paleontology, and biogeography. He worked extensively in cata ...
in 1893. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen of ''Argyrosaurus superbus'' is a huge left forelimb, MLP 77-V-29-1. found at Chico River, a
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
/
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
horizon in the Lago Colhué Huapi Formation. The material includes the
humerus The humerus (; : humeri) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius (bone), radius and ulna, and consists of three sections. The humeral upper extrem ...
,
ulna The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...
,
radius In classical geometry, a radius (: radii or radiuses) of a circle or sphere is any of the line segments from its Centre (geometry), center to its perimeter, and in more modern usage, it is also their length. The radius of a regular polygon is th ...
, and all five
metacarpals In human anatomy, the metacarpal bones or metacarpus, also known as the "palm bones", are the appendicular skeleton, appendicular bones that form the intermediate part of the hand between the phalanges (fingers) and the carpal bones (wrist, wris ...
. Recent stratigraphic information from the Lago Colhue Huapi Formation has the area that the holotype was collected from being a more probable early late Maastrichtian age instead of early Coniacian. When discovered, the forelimb was apparently a part of a complete skeleton, however, during excavation the rest of the remains were completely destroyed. In the same publication, Lydekker assigned a partial left femur from
Chubut Province Chubut ( from Tehuelche language, Tehuelche 'transparent'; ) is a provinces of Argentina, province in southern Argentina, situated between the 42nd parallel south (the border with Río Negro Province), the 46th parallel south (bordering Santa ...
, MLP 21, to ''Argyrosaurus'' but he did not explain his reasoning why he thought the femur was referable to the genus. He also assigned two
caudal vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...
centra Centra is a convenience shop chain that operates throughout Ireland. The chain operates as a symbol group owned by Musgrave Group, the food wholesaler, meaning the individual shops are all owned by individual franchisees. The chain has three ...
, MLP 22, from the Santa Cruz Province. After these specimens, other remains were found, including other limb bones, a clavicle, a pubic bone and some tail vertebrae that were referred to the same genus. In 1929
Friedrich von Huene Baron Friedrich Richard von Hoyningen-Huene (22 March 1875 – 4 April 1969) was a German nobleman paleontologist who described a large number of dinosaurs, more than anyone else in 20th-century Europe. He studied a range of Permo-Carbonife ...
referred several specimens to ''Argyrosaurus''; a caudal centrum (DGM
umber unknown Umber is a natural earth pigment consisting of iron oxide and manganese oxide; it has a brownish color that can vary among shades of yellow, red, and green. Umber is considered one of the oldest pigments known to humans, first used in the Ajant ...
from Chubut Province, three caudal centra MACN 5205 from Santa Cruz Province, a right femur MLP 27 from
Neuquén Province Neuquén () is a Provinces of Argentina, province of Argentina, located in the west of the country, at the northern end of Patagonia. It borders Mendoza Province to the north, Rio Negro Province to the southeast, and Chile to the west. It also me ...
, another right femur FMNH 13018 which measures over long from Chubut Province, a small right humerus MACN 5017 from Neuquén Province, and the distal end of a humerus, half of a radius, and a rib fragment, MMAB
umber unknown Umber is a natural earth pigment consisting of iron oxide and manganese oxide; it has a brownish color that can vary among shades of yellow, red, and green. Umber is considered one of the oldest pigments known to humans, first used in the Ajant ...
from
Uruguay Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...
. A complete right femur FMNH 13019 and left tibia FMNH 13020 from Chubut Province were originally referred to ''
Antarctosaurus ''Antarctosaurus'' (; meaning "southern lizard") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period (geology), Period of what is now South America. The type species, ''Antarctosaurus wichmannianus'', and a second speci ...
'' by Von Huene in 1929, however, in 2003 Jaime Eduardo Powell tentatively referred them to cf. ''Argyrosaurus''. In 1979 Bonaparte and Gasparini referred a partial skeleton, PLV 4628/MACN-CH 217, which included limb material and several vertebrae from Chubut Province to ''Argyrosaurus''. Although these numerous other remains have been referred, in 2012 Philip Mannion and Alejandro Otero considered the holotype remains the only material which unambiguously pertains to the genus. Many of the referred specimens either have no overlapping material, lack key features in the holotype, or were not illustrated making not possible to assign them without being restudied. Most of the referred specimens were considered indeterminate titanosaurs. They also considered one of the referred specimens, the partial skeleton PLV 4628/MACN-CH 217, to be its own genus which they called ''
Elaltitan ''Elaltitan'' is an extinct genus of large lithostrotian titanosaur sauropod dinosaur known from the Late Cretaceous (mid Cenomanian to Turonian stage) of Chubut Province, southern Argentina. It contains a single species, ''Elaltitan lilloi''. ...
''. The genus name means 'silver lizard' from
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
''argyros'', 'silver', and ''sauros'', 'lizard', because it was discovered in
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
, which literally means 'silver land'. The
specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
means "proud" in
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
.


Description

''Argyrosaurus'' was a medium-sized sauropod which is estimated to measure long and weighing up to according to Paul.Paul, Gregory S. ''Dinosaurs: A Field Guide''. London: A. & C. Black, 2010. Print. However, in 2012 Thomas Holtz gave a higher estimation at in length and a possible weight of 43.5-50.8 tonnes (48-56 short tons). More recently it was listed at and 26 tonnes (28.6 short tons). It is distinguished from other genera by the following unique features: * the medial side of the humerus has a wide ridge that projects forward * the midshaft of the humerus is extremely compressed * the width of the distal end of the radius is only slightly greater than the midshaft width * the radius is subtriangular when viewed from the distal end * the metacarpals are extremely elongated


References


External links


An image of ''Argyrosaurus''

Artist's impression of ''Argyrosaurus'', with scaled holotype forelimb
{{Portal bar, Dinosaurs, Argentina Lithostrotia Dinosaur genera Maastrichtian dinosaurs Dinosaurs of Argentina Fossil taxa described in 1893 Taxa named by Richard Lydekker