Argentine Espinal
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The Espinal (NT0801) is an
ecoregion An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and c ...
of dry, thorny forest, savanna and steppe in Argentina. It has been extensively modified by large scale cattle ranching, but remnants of the original flora remain. It is threatened by the advance of the irrigation-based agricultural frontier.


Location

The espinal is a broad arc of arid land in the interior of Argentina to the north and west of
Buenos Aires Buenos Aires, controlled by the government of the Autonomous City of Buenos Aires, is the Capital city, capital and largest city of Argentina. It is located on the southwest of the Río de la Plata. Buenos Aires is classified as an Alpha− glob ...
, extending from the northern Uruguay border west through Santa Fe to Cordoba and south to the
Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...
coast south of
Bahía Blanca Bahía Blanca (; English: ''White Bay''), colloquially referred to by its own local inhabitants as simply Bahía, is a city in the Buenos Aires Province, Buenos Aires province of Argentina, centered on the northwestern end of the eponymous Blanc ...
. It includes central
Santa Fe Province The Invincible Province of Santa Fe (, , lit. "Holy Faith") is a Provinces of Argentina, province of Argentina, located in the center-east of the country. Neighboring provinces are from the north clockwise Chaco Province, Chaco (divided by the 2 ...
, Córdoba Province and the northern part of
San Luis Province San Luis () is a Provinces of Argentina, province of Argentina located near the geographical center of the country (on the 32° South parallel). Neighboring provinces are, from the north clockwise, La Rioja Province (Argentina), La Rioja, Córdo ...
. It has an area of . The name "''espinal''" (thorny) refers to the thorny deciduous shrubland forest of the region between the flooded wetlands along the
Paraná River The Paraná River ( ; ; ) is a river in south-central South America, running through Brazil, Paraguay, and Argentina for some ."Parana River". Encyclopædia Britannica. Encyclopædia Britannica Online. Encyclopædia Britannica Inc., 2012. Web. ...
to the east and the
Sierras de Córdoba The Sierras de Córdoba is a mountain range in central Argentina, located between the Pampas to the east and south and the Gran Chaco, Chaco to the north and east. Most of the range is located in Córdoba Province, Argentina, Córdoba Province ...
to the west. The northern part of the ecoregion adjoins the Humid Pampas around Buenos Aires to the south, the Uruguayan savanna to the east, and the
Humid Chaco The Humid Chaco ( Spanish: ''Chaco Húmedo'' or ''Chaco Oriental'') is a tropical grasslands, savannas, and shrublands ecoregion in South America. It lies in the basin of the Paraná River, covering portions of central Paraguay and northern Arge ...
and Dry Chaco to the north and northwest. A belt of Paraná flooded savanna runs from north to south through the ecoregion along the Paraná River. The southern part of the ecoregion lies between the Low Monte ecoregion to the west and the Humid Pampas to the east.


Physical

The region mostly covers flat plains with
loess A loess (, ; from ) is a clastic rock, clastic, predominantly silt-sized sediment that is formed by the accumulation of wind-blown dust. Ten percent of Earth's land area is covered by loesses or similar deposition (geology), deposits. A loess ...
or sandy soils. There are wide variations in climate. The climate in the north is warm and wet, with rains in the summer. The
Köppen climate classification The Köppen climate classification divides Earth climates into five main climate groups, with each group being divided based on patterns of seasonal precipitation and temperature. The five main groups are ''A'' (tropical), ''B'' (arid), ''C'' (te ...
is "Cfa": warm temperate, fully humid, hot summer. At a sample location at coordinates the mean temperature ranges from in July to in January. The yearly average mean temperature is about . Yearly total precipitation is about . Monthly precipitation ranges from in June to in March.


Ecology

The espinal is in the
Neotropical realm The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropics, tropical Ecoregion#Terrestrial, terrestrial ecoregions of the Americas and the entire South American temperat ...
, in the
temperate grasslands, savannas, and shrublands Temperate grasslands, savannas, and shrublands are terrestrial biomes defined by the World Wide Fund for Nature. The predominant vegetation in these biomes consists of grass and/or shrubs. The climate is temperate and ranges from Semi-arid clima ...
biome. The dry ecoregion was once home to many species of birds, mammals and plants. The introduction of cattle in the 17th century had a huge impact, and the original habitat is now found only in isolated patches.


Flora

Vegetation includes deciduous
xerophytic A xerophyte () is a species of plant that has adaptations to survive in an environment with little liquid water. Examples of xerophytes include cactus, cacti, pineapple and some gymnosperm plants. The morphology (biology), morphology and physiology ...
forests, palm groves, grassy savannahs, grassy steppes and bushy steppes. Plants of the
Prosopis ''Prosopis'' is a genus of flowering plants in the family Fabaceae. The current circumscription of the genus contains three species found in northern Africa, the Middle East, Central and South Asia. Previously it also contained around 40 species ...
and
Acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...
genera of the
Mimosoideae The Mimosoideae are a traditional subfamily of trees, herbs, lianas, and shrubs in the pea family (Fabaceae) that mostly grow in tropical and subtropical climates. They are typically characterized by having radially symmetric flowers, with petal ...
family are common, add the mature forest areas are dominated by trees of the Prosopis genus. Other plants belong to genera such as Jodina (
Santalaceae The Santalaceae, sandalwoods, are a widely distributed family of flowering plants (including small trees, shrubs, perennial plants, perennial herbs, and epiphyte, epiphytic climbersHewson & George t al.'Santalaceae'' taxonomy, 1984, pp. 191-1 ...
family),
Celtis ''Celtis'' is a genus of about 60–70 species of deciduous trees, commonly known as hackberries or nettle trees, in the hemp family Cannabaceae. It has a cosmopolitan distribution. Description ''Celtis'' species are generally medium-sized tree ...
, Schinus, Geoffroea and Atamisquea (
Capparaceae The Capparaceae (or Capparidaceae), commonly known as the caper family, are a family of plants in the order Brassicales. As currently circumscribed, the family contains 15 genera and about 430 species. The largest genera are ''Capparis'' (about 1 ...
family). In the ''carob'' subregion in the north of the espinal the main tree species include ''
Prosopis nigra ''Neltuma nigra'' (formerly ''Prosopis nigra'') is a South American leguminous tree species that inhabits the Gran Chaco ecoregion (in particular, the transition zone between the Wet Chaco and the Southern Chaco), in Argentina and Paraguay. It ...
'', ''
Prosopis alba ''Neltuma alba'' (formerly ''Prosopis alba'') is a South American tree species that grows in central Argentina, the Gran Chaco ecoregion, and part of the Argentine Mesopotamia, as well as Bolivia, Chile, Paraguay, and Peru. It is known as ''alg ...
'', '' Celtis tala'', ''
Geoffroea decorticans ''Geoffroea decorticans'', the chañar, kumbaru, or Chilean palo verde (green wood), is a small deciduous tree, up to 8 meters (25 ft) tall that inhabits most arid forests (montes or espinales) of southern South America. The chañar is cold ...
'', '' Vachellia caven'', '' Jodina rhombifolia'', '' Scutia buxifolia'' and '' Schinus longifolia''. Isolated specimens or groups of the '' Trithrinax campestris'' palm are still found in some areas. The "caldén" subregion in the south of the espinal has xerophytic forests with many '' Prosopis caldenia'' (calden) trees, an endemic species. It also contains areas of grassy savanna, dunes with flora adapted to sand, and areas with salty soils holding scrublands or
halophilic A halophile (from the Greek word for 'salt-loving') is an extremophile that thrives in high salt concentrations. In chemical terms, halophile refers to a Lewis acidic species that has some ability to extract halides from other chemical species. ...
steppes.


Fauna

There are three species of endemic birds. The Mar Chiquita lake and the Dulce River swamps are designated a
Ramsar wetland The Ramsar Convention on Wetlands of International Importance Especially as Waterfowl Habitat is an international treaty for the conservation and sustainable use of Ramsar sites (wetlands). It is also known as the Convention on Wetlands. It i ...
that is a key site for endangered neotropical birds. The wetland supports a rich and diverse fauna with 138 known species of birds, including large breeding groups of
Chilean flamingo The Chilean flamingo (''Phoenicopterus chilensis'') is a species of large flamingo at a height of closely related to the American flamingo and the greater flamingo, with which it was previously considered a subspecies before being classified ...
(''Phoenicopterus chilensis''). Native species include
guanaco The guanaco ( ; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids; the other species is the vicuña, which lives at higher elevations. Etymology The gua ...
s,
greater rhea The greater rhea (''Rhea americana'') is a species of flightless bird native to eastern South America. Other names for the greater rhea include the grey, common, or American rhea; ema (Portuguese (language), Portuguese); or ñandú (Guaraní lang ...
s, and
giant anteater The giant anteater (''Myrmecophaga tridactyla'') is an Insectivore, insectivorous mammal native to Central America, Central and South America. It is the largest of the four living species of anteaters, which are classified with sloths in the or ...
s. Endangered amphibians include blunt-headed salamander (''Ambystoma amblycephalum''). Endangered birds include yellow cardinal (''Gubernatrix cristata''), Chaco eagle (''Buteogallus coronatus'') and Eskimo curlew (''Numenius borealis'').


Status

The
World Wildlife Fund The World Wide Fund for Nature (WWF) is a Swiss-based international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the ...
gives the espinal ecoregion the status of "Critical/Endangered". Much of the ecoregion has been used for agriculture, and the trees of the sparse dry forests have been used for firewood, fence posts, flooring and furniture. Irrigation systems are expanding the agricultural frontier into area that were previously not affected. The Dulce River floodplain and Mar Chiquita lake are threatened by increasing use of the water upstream. In parts of the ecoregion, woody plant encroachment has been observed.


Protected areas

1.74% of the ecoregion is in protected areas. They include the Lihué Calel National Park, La Reforma University Reserve, Chacharramendi Provincial Reserve and Luro Provincial Reserve. Other units are the
El Palmar National Park El Palmar National Park (in Spanish, ''Parque Nacional El Palmar'') is one of Argentina's national parks. It is located on the center-east of the province of Entre Ríos, midway between the cities of Colón (54 km) and Concordia (60  ...
and the Parque Atlántico Mar Chiquita.


References


Sources

* * * {{DEFAULTSORT:Espinal Neotropical ecoregions Ecoregions of Argentina Temperate grasslands, savannas, and shrublands