Turnstones are two

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

that constitute the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

''Arenaria'' in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Scolopacidae Scolopacidae is a large family of shorebirds, or waders, which mainly includes many species known as sandpipers, but also others such as woodcocks, curlews and snipes. Most of these species eat small invertebrates picked out of the mud or soil ...

. They are closely related to

calidrid ''Calidris'' is a genus of Arctic-breeding, strongly migratory wading birds in the family Scolopacidae. These birds form huge mixed flocks on coasts and estuaries in winter. Migratory shorebirds are shown to have decline in reproductive traits ...

sandpipers and might be considered members of the

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...

Calidriini. The

''Arenaria'' was introduced by the French zoologist

Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...

in 1760 with the

ruddy turnstone The ruddy turnstone (''Arenaria interpres'') is a small Cosmopolitan distribution, cosmopolitan wader, wading bird, one of two species of turnstone in the genus ''Arenaria''. It is now classified in the sandpiper family Scolopacidae but was form ...

(''Arenaria interpres'') as the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

. The genus name ''arenaria'' is from

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

''arenarius'', "inhabiting sand", from ''arena'', "sand". The genus contains two species: the

(''Arenaria interpres'') and the

black turnstone The black turnstone (''Arenaria melanocephala'') is a species of small wading bird. It is one of two species of turnstone in the genus ''Arenaria'' the ruddy turnstone (''A. interpres'') being the other. It is now classified in the sandpiper fam ...

(''Arenaria melanocephala''). Both birds are

wader 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...

s. Their length is typically between 20 and 25 cm, with a wingspan between 50 and 60 cm and a body mass between 110 and 130g. For waders their build is stocky, with short, slightly upturned, wedge shaped bills. They have white patches on the back, wings and tail. They are high

Arctic The Arctic (; . ) is the polar regions of Earth, polar region of Earth that surrounds the North Pole, lying within the Arctic Circle. The Arctic region, from the IERS Reference Meridian travelling east, consists of parts of northern Norway ( ...

breeders, and are migratory. Their strong necks and powerful, slightly upturned bills are adapted to their feeding technique. As the name implies, these species overturn stones, seaweed, and similar items in search of

invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...

prey.Svensson, Lars et al. Collins Bird Guide 2nd ed. Publisher: Collins 2010. They are strictly coastal, prefer stony beaches to sand, and often share beach space with other species of waders such as

purple sandpiper The purple sandpiper (''Calidris maritima'') is a small shorebird in the sandpiper family Scolopacidae. This is a hardy sandpiper that breeds in the arctic and subarctic regions of Eurasia and North America and winters further south on the Atlan ...

Species

There exists a

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

bone, a

distal Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provi ...

piece of

tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bird bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) a ...

found in the

Edson Beds Edson may refer to: Places Canada * Edson, Alberta United States * Edson, Kansas, an unincorporated community * Edson, South Dakota, a ghost town * Edson, Wisconsin, a town ** Edson (community), Wisconsin, an unincorporated community People Giv ...

Sherman County, Kansas Sherman County is a county located in the U.S. state of Kansas. Its county seat is Goodland. As of the 2020 census, the county population was 5,927. Sherman County was created by the Legislature of 1873, and named after general William Sh ...

. Dating from the mid-

Blancan The Blancan North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 4,750,000 to 1,806,000 years BP, a period of .pectoral sandpiper The pectoral sandpiper (''Calidris melanotos'') (often abbreviated pec) is a small, Bird migration, migratory wader that breeds in North America and Palearctic, Asia, wintering in South America and Oceania. It eats small invertebrates. Its bird ...

, but has some traits reminiscent of turnstones. Depending on which traits are

apomorph In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to ha ...

ic and

plesiomorph In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral Phenotypic trait, character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorph ...

ic, it may be an ancestral representative of either lineage.

References

External links

* Ruddy turnstone = turnstone
Species text in ''The Atlas of Southern African Birds''

– Cornell Lab of Ornithology

– USGS Patuxent Bird Identification InfoCenter
Turnstones feeding and bathing
{{Taxonbar, from=Q4865776 Sandpipers Taxa named by Mathurin Jacques Brisson