''Archaeothyris'' is an extinct
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
ophiacodontid
Ophiacodontidae is an extinct family of early eupelycosaurs from the Carboniferous and Permian. ''Archaeothyris'', and ''Clepsydrops'' were among the earliest ophiacodontids, appearing in the Late Carboniferous. Ophiacodontids are among the most ...
synapsid
Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes rep ...
that lived during the
Late Carboniferous
Late may refer to:
* LATE, an acronym which could stand for:
** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia
** Local-authority trading enterprise, a New Zealand business law
** Local average treatment effe ...
and is known from
Nova Scotia
Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland".
Most of the population are native En ...
. Dated to 306 million years ago, ''Archaeothyris'', along with a more poorly known synapsid called ''
Echinerpeton'', are the oldest undisputed synapsids known. The name means ''ancient window'' (Greek), and refers to the opening in the skull, the
temporal fenestra, which indicates this is an early synapsid. ''
Protoclepsydrops'' also from Nova Scotia is slightly older but is known by very fragmentary materials.
Description

''Archaeothyris'' was more advanced than the early
sauropsids, having strong jaws that could open wider than those of the early
reptiles. While its sharp teeth were all of the same size & shape, it did possess a pair of enlarged
canine
Canine may refer to:
Zoology and anatomy
* a dog-like Canid animal in the subfamily Caninae
** '' Canis'', a genus including dogs, wolves, coyotes, and jackals
** Dog, the domestic dog
* Canine tooth, in mammalian oral anatomy
People with the ...
s, suggesting that it was a carnivore.
Archaeothyris' legs were articulated laterally at its pelvis and shoulders, which gave it a sprawling stance. The first toe is smaller than the second.
Classification
''Archaeothyris'' belonged to the family
Ophiacodontidae
Ophiacodontidae is an extinct family of early eupelycosaurs from the Carboniferous and Permian. '' Archaeothyris'', and ''Clepsydrops'' were among the earliest ophiacodontids, appearing in the Late Carboniferous. Ophiacodontids are among the m ...
, a group of early pelycosaurs that evolved early in the
Late Carboniferous
Late may refer to:
* LATE, an acronym which could stand for:
** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia
** Local-authority trading enterprise, a New Zealand business law
** Local average treatment effe ...
. It was one of the earliest and most basal synapsids (the group which includes
mammals).
Below is a
cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
modified from the analysis of Benson (2012):
Discovery and paleoecology
Fossils of ''Archaeothyris'' were first described in 1972 from the Joggins fossil cliffs, the same locality in which the early reptiles ''
Hylonomus'' and ''
Petrolacosaurus
''Petrolacosaurus'' ("rock lake lizard") is an extinct genus of diapsid reptile from the late Carboniferous period. It was a small, long reptile, and the earliest known reptile with two temporal fenestrae (holes at the rear part of the skull) ...
'' (both of which resemble ''Archaeothyris'') were found.
''Archaeothyris'' lived in what is now
Nova Scotia
Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland".
Most of the population are native En ...
, about 306 million years ago in the Carboniferous Period (
Pennsylvanian Pennsylvanian may refer to:
* A person or thing from Pennsylvania
* Pennsylvanian (geology)
The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, in the International Commission on Stratigraphy, ICS geologic timesca ...
).
[Hess J.C., Lippolt H.J. (1986): 40Ar/39Ar ages of tonstein and tuff sanidines: new calibration points for the improvement of the Upper Carboniferous time scale. ''Chem Geol'' no 59: pp 143–154]
Nova Scotia at this time was a swamp, similar to today's
Everglades
The Everglades is a natural region
A natural region (landscape unit) is a basic geographic unit. Usually, it is a region which is distinguished by its common natural features of geography, geology, and climate.
From the ecological point o ...
in
Florida
Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, a ...
. The "trees" (actually giant
club mosses) were very tall, some, such as ''
Lepidodendron
''Lepidodendron'' is an extinct genus of primitive vascular plants belonging to the family Lepidodendraceae, part of a group of Lycopodiopsida known as scale trees or arborescent lycophytes, related to quillworts and lycopsids (club mosses). T ...
'', up to tall. ''Archaeothyris'' and the other early amniotes lived in the moist vegetation on the forest ground, together with the more terrestrially adapted
labyrinthodont
"Labyrinthodontia" (Greek, 'maze-toothed') is an informal grouping of extinct predatory amphibians which were major components of ecosystems in the late Paleozoic and early Mesozoic eras (about 390 to 150 million years ago). Traditionally conside ...
amphibians.
See also
*
List of pelycosaurs
This list of pelycosaurs is an attempt to create a comprehensive listing of all genera that have ever been included in the synapsida excluding therapsida and purely vernacular terms. The list includes all commonly accepted genera, but also genera t ...
*
Evolution of mammals
The evolution of mammals has passed through many stages since the first appearance of their synapsid ancestors in the Pennsylvanian (geology), Pennsylvanian sub-period of the late Carboniferous period. By the mid-Triassic, there were many synaps ...
*
List of transitional fossils
This is a partial list of transitional fossils (fossil remains of groups that exhibit both "primitive" and derived traits). The fossils are listed in series, showing the transition from one group to another, representing significant steps in ...
*
Carboniferous tetrapods
References
Further reading
*
External links
Transitional Vertebrate Fossils- includes description of important
transitional genera from reptile to mammal (includes a little information about ''Archaeothyris'')
{{Taxonbar, from=Q133034
Ophiacodontids
Prehistoric synapsid genera
Carboniferous synapsids
Carboniferous synapsids of North America
Transitional fossils
Taxa named by Robert R. Reisz
Fossil taxa described in 1972
Paleozoic life of Nova Scotia