''Aquilegia eximia'', the serpentine columbine or Van Houtte's columbine, is a
perennial
In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
species of
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the family
Ranunculaceae
Ranunculaceae (, buttercup or crowfoot family; Latin "little frog", from "frog") is a family (biology), family of over 2,000 known species of flowering plants in 43 genera, distributed worldwide.
The largest genera are ''Ranunculus'' (600 spec ...
,
endemic
Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
California
California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
.
Description
''Aquilegia eximia'' is a perennial
herb
Herbs are a widely distributed and widespread group of plants, excluding vegetables, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal purposes, or for fragrances. Culinary use typically distingu ...
growing from a thick
caudex
A caudex (: caudices) of a plant is a stem, but the term is also used to mean a rootstock and particularly a basal stem structure from which new growth arises.pages 456 and 695
In the strict sense of the term, meaning a stem, "caudex" is most ...
and reaching a maximum height of . The lower leaves are divided into large, leaflike segments up to long and a lobed oval. Leaves higher up the stem are not segmented, but may be deeply lobed. The
inflorescence
In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
bears a large, nodding columbine flower. Each flower has five bright red to orange-red flat
sepal
A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106
Etymology
The term ''sepalum'' ...
s of in length, and five petals which have hollow spurs long, bright orange-red on the outer surface and lighter orange to yellow inside. The mouth of each hollow petal tube is wide. The sepals and petals are generally reflexed back toward the stem and the five
pistils
Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
and many thin
stamen
The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10
Morphology and terminology
A stamen typically consists of a stalk called the filament ...
s extend forward from the center of the flower.
Taxonomy
The species is part of a
clade
In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
containing all the
North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
n species of columbines, that likely split from their closest relatives in
East Asia
East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...
in the mid-
Pliocene
The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58
Etymology
The
specific epithet
In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
'' eximia'' means "excellent in size or beauty, choice, distinguished" in Latin.
Distribution and habitat
''Aquilegia eximia'' is endemic to the mixed-evergreen or conifer forests of the
California Coast Ranges
The Coast Ranges of California span from Del Norte County, California, Del Norte or Humboldt County, California, south to Santa Barbara County, California, Santa Barbara County. The other three coastal California mountain ranges are the Trans ...
, the western
Transverse Ranges
The Transverse Ranges are a group of mountain ranges of Southern California, in the Pacific Coast Ranges physiographic region in North America. The Transverse Ranges begin at the southern end of the California Coast Ranges and lie within Santa Ba ...
, and the
San Francisco Bay Area
The San Francisco Bay Area, commonly known as the Bay Area, is a List of regions of California, region of California surrounding and including San Francisco Bay, and anchored by the cities of Oakland, San Francisco, and San Jose, California, S ...
, usually growing on
serpentine soil
Serpentine soil is an uncommon soil type produced by weathered ultramafic rock such as peridotite and its metamorphic derivatives such as serpentinite. More precisely, serpentine soil contains minerals of the serpentine subgroup, especially an ...
s and sometimes in moist ravines at altitudes of .
Ecology
The species flowers from May to October and is pollinated by
hummingbird
Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
s. Its range overlaps with that of ''
Aquilegia formosa
''Aquilegia formosa'', the crimson columbine, western columbine, or (ambiguously) "red columbine", is a common wildflower native to western North America, from Alaska to Baja California, and eastward to Montana and Wyoming.
Description
''Aquileg ...
'', which flowers earlier in the year.
Conservation
,
NatureServe
NatureServe, Inc. is a non-profit organization based in Arlington County, Virginia, United States, US, that provides proprietary wildlife conservation-related data, tools, and services to private and government clients, partner organizations, and ...
listed ''Aquilegia eximia'' as Vulnerable (G3). This status is noted as being in need of review, having last been assessed on .