The Barbary sheep (''Ammotragus lervia''), also known as aoudad (pronounced �ɑʊdæd is a

species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

of caprine native to rocky mountains in

North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in t ...

. While this is the only species in genus ''Ammotragus'', six subspecies have been described. Although it is rare in its native North Africa, it has been introduced to North America, southern

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...

, and elsewhere. It is also known in the

Berber language The Berber languages, also known as the Amazigh languages or Tamazight,, ber, label= Tuareg Tifinagh, ⵜⵎⵣⵗⵜ, ) are a branch of the Afroasiatic language family. They comprise a group of closely related languages spoken by Berber comm ...

as ''waddan'' or ''arwi'', and in former French territories as the moufflon.

Description

Barbary sheep stand tall at the shoulder, with a length around , and weigh

They are sandy-brown, darkening with age, with a slightly lighter underbelly and a darker line along the back. Upper parts and the outer parts of the legs are a uniform reddish- or grayish-brown. Some shaggy hair is on the

throat In vertebrate anatomy, the throat is the front part of the neck, internally positioned in front of the vertebrae. It contains the pharynx and larynx. An important section of it is the epiglottis, separating the esophagus from the trachea (windpi ...

(extending down to the chest in males) with a sparse

mane Mane may refer to: * Mane (horse), the line of hair along the spine of the neck * Mane (lion), the hair found around the male mammal's neck In arts and entertainment * ''Mane'' (film) is a 1990 Kannada language film directed by Girish Kasaravall ...

. Their horns have a

triangular A triangle is a polygon with three edges and three vertices. It is one of the basic shapes in geometry. A triangle with vertices ''A'', ''B'', and ''C'' is denoted \triangle ABC. In Euclidean geometry, any three points, when non-collinear, ...

cross-section Cross section may refer to: * Cross section (geometry) ** Cross-sectional views in architecture & engineering 3D *Cross section (geology) * Cross section (electronics) * Radar cross section, measure of detectability * Cross section (physics) **Ab ...

. The horns curve outward, backward, then inward, and can exceed in length. The horns are fairly smooth, with slight wrinkles evident at the base as the animal matures.

Range

Natural range

Barbary sheep naturally occur in northern Africa in

Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...

Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , religi ...

Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...

, northern

Chad Chad (; ar, تشاد , ; french: Tchad, ), officially the Republic of Chad, '; ) is a landlocked country at the crossroads of North and Central Africa. It is bordered by Libya to the north, Sudan to the east, the Central African Repub ...

Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Med ...

Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to the east, Su ...

, northern

Mali Mali (; ), officially the Republic of Mali,, , ff, 𞤈𞤫𞤲𞥆𞤣𞤢𞥄𞤲𞤣𞤭 𞤃𞤢𞥄𞤤𞤭, Renndaandi Maali, italics=no, ar, جمهورية مالي, Jumhūriyyāt Mālī is a landlocked country in West Africa. Ma ...

, Mauritania,

Niger ) , official_languages = , languages_type = National languagesSudan (west of the

Nile The Nile, , Bohairic , lg, Kiira , Nobiin language, Nobiin: Áman Dawū is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa and has historically been considered ...

, and in the Red Sea Hills east of the Nile).

Introduced populations

Barbary sheep have been introduced to southeastern

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' ( Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...

, the southwestern

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...

( Chinati Mountains on La Escalera Ranch,

Guadalupe Mountains National Park Guadalupe Mountains National Park is an American national park in the Guadalupe Mountains, east of El Paso, Texas. The mountain range includes Guadalupe Peak, the highest point in Texas at , and El Capitan used as a landmark by travelers on the ...

Palo Duro Canyon Palo Duro Canyon is a canyon system of the Caprock Escarpment located in the Texas Panhandle near the cities of Amarillo and Canyon. As the second-largest canyon in the United States, it is roughly long and has an average width of , but reaches ...

, the Trans-Pecos, and other parts of

Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...

and

New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...

Niihau Niihau ( Hawaiian: ), anglicized as Niihau ( ), is the westernmost main and seventh largest inhabited island in Hawaii. It is southwest of Kauaʻi across the Kaulakahi Channel. Its area is . Several intermittent playa lakes provide wetland habi ...

Island (Hawaii),

Mexico Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...

, and some parts of Africa. They have become common in a limited region of southeastern

, since its introduction in 1970 to

Sierra Espuña The Sierra Espuña is a mountain range in the Region of Murcia, Spain. It is part of the Penibaetic System. The Sierra Espuña Regional Park protects 17,804 ha of the mountain range in the municipalities of Alhama de Murcia, Totana and Mula. The ...

regional park A regional park is an area of land preserved on account of its natural beauty, historic interest, recreational use or other reason, and under the administration of a form of local government. Definition A regional park can be a special park distri ...

as a

game A game is a structured form of play, usually undertaken for entertainment or fun, and sometimes used as an educational tool. Many games are also considered to be work (such as professional players of spectator sports or games) or art (su ...

species. Its adaptability enabled it to colonise nearby areas quickly, and private game estates provided other centers of dispersion. The species is currently expanding, according to recent field surveys, now being found in the provinces of

Alicante Alicante ( ca-valencia, Alacant) is a city and municipality in the Valencian Community, Spain. It is the capital of the province of Alicante and a historic Mediterranean port. The population of the city was 337,482 , the second-largest in th ...

Almería Almería (, , ) is a city and municipality of Spain, located in Andalusia. It is the capital of the province of the same name. It lies on southeastern Iberia on the Mediterranean Sea. Caliph Abd al-Rahman III founded the city in 955. The city gr ...

Granada Granada (,, DIN: ; grc, Ἐλιβύργη, Elibýrgē; la, Illiberis or . ) is the capital city of the province of Granada, in the autonomous community of Andalusia, Spain. Granada is located at the foot of the Sierra Nevada mountains, at the c ...

, and

Murcia Murcia (, , ) is a city in south-eastern Spain, the Capital (political), capital and most populous city of the autonomous community of the Region of Murcia, and the List of municipalities of Spain, seventh largest city in the country. It has a ...

. This species is a potential competitor to native

ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraff ...

s inhabiting the

Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...

. The species has also been introduced to

La Palma La Palma (, ), also known as ''La isla bonita'' () and officially San Miguel de La Palma, is the most north-westerly island of the Canary Islands, Spain. La Palma has an area of making it the fifth largest of the eight main Canary Islands. The ...

(

Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Mo ...

), and has spread throughout the northern and central parts of the island, where it is a serious threat to

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...

vegetation Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic charact ...

. Aoudad has also been introduced in

Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...

several times and there is a population in

Mosor Mosor ( la, Massarus), or Mount Mosor (''Massarus Mons''; it, Monte Massaro), is a mountain range in Croatia located near the city of Split on the Adriatic coast. It belongs to Dinaric Alps, and it stretches from the pass of Klis in the north ...

Taxonomy

''A. lervia'' is the only species in the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

''Ammotragus''. However, some authors include this genus in the goat genus '' Capra'', together with the sheep genus ''

Ovis ''Ovis'' is a genus of mammals, part of the Caprinae subfamily of the ruminant family Bovidae. Its seven highly sociable species are known as sheep or ovines. Domestic sheep are members of the genus, and are thought to be descended from the w ...

''. The subspecies are found

allopatric Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...

ally in various parts of North Africa: * ''A. l. lervia'' Pallas, 1777 (vulnerable) * ''A. l. ornata'' I. Geoffroy Saint-Hilaire, 1827 (Egyptian Barbary sheep, thought to be extinct in the wild but still found in the eastern desert of Egypt)Wacher, T., El Din, S. B., Mikhail, G., & El Din, M. B. (2002). New observations of the ‘extinct’ Barbary sheep ''Ammotragus lervia ornata'' in Egypt. ''Oryx'', 36(3), 301-304. Manlius, N., Menardi-Noguera, A. and Zboray, A. 2003. Decline of the Barbary sheep (''Ammotragus lervia'') in Egypt during the 20th century: literature review and recent observations. ''Journal of Zoology'' (London) 259: 403-409. * ''A. l. sahariensis'' Rothschild, 1913 (vulnerable) * ''A. l. blainei'' Rothschild, 1913 (vulnerable) * ''A. l. angusi'' Rothschild, 1921 (vulnerable) * ''A. l. fassini'' Lepri, 1930 (vulnerable)

Habitats

Barbary sheep are found in

arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ...

mountainous areas where they graze and browse

grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in law ...

es,

bushes A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

, and lichens. They are able to obtain all their

metabolic water Metabolic water refers to water created inside a living organism through their metabolism, by oxidizing energy-containing substances in their food. Animal metabolism produces about 107-110 grams of water per 100 grams of fat, 41-42 grams of water ...

from food, but if liquid water is available, they drink and wallow in it. Barbary sheep are

crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of dayli ...

- active in the early morning and late afternoon and rest in the heat of the day. They are very agile and can achieve a standing jump over . They are well adapted to their habitat, which consist of steep, rocky mountains and canyons. They often flee at the first sign of danger, typically running uphill. They are extremely nomadic and travel constantly via mountain ranges. Their main

predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...

s in North Africa were the

Barbary leopard The African leopard (''Panthera pardus pardus'') is the nominate subspecies of the leopard, native to many countries in Africa. It is widely distributed in most of sub-Saharan Africa, but the historical range has been fragmented in the course of ...

, the

Barbary lion The Barbary lion, also called the North African lion, Berber lion, Atlas lion, and Egyptian lion, is an extinct population of the lion subspecies ''Panthera leo leo''. It lived in the mountains and deserts of the Barbary Coast of North Africa, f ...

, and

caracal The caracal (''Caracal caracal'') () is a medium-sized wild cat native to Africa, the Middle East, Central Asia, and arid areas of Pakistan and northwestern India. It is characterised by a robust build, long legs, a short face, long tufted e ...

, but now humans, feral dogs, competition due to overgrazing by domestic animals and drought threaten their populations.

Names

The

binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''Ammotragus lervia'' derives from the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

ἄμμος ''ámmos'' ("sand", referring to the sand-coloured coat) and τράγος ''trágos'' ("goat"). ''Lervia'' derives from the wild sheep of northern Africa described as "lerwee" by Rev. T. Shaw in his "Travels and Observations" about parts of Barbary and

Levant The Levant () is an approximation, approximate historical geography, historical geographical term referring to a large area in the Eastern Mediterranean region of Western Asia. In its narrowest sense, which is in use today in archaeology an ...

. The

Spanish Spanish might refer to: * Items from or related to Spain: ** Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Ca ...

named this sheep the ''arruis'', from Berber ''arrwis'', and the

Spanish Legion For centuries, Spain recruited foreign soldiers to its army, forming the Foreign Regiments () - such as the Regiment of Hibernia (formed in 1709 from Irishmen who fled their own country in the wake of the Flight of the Earls and the pen ...

even used it as a mascot for a time. ''Aoudad'' () is the name for this sheep used by the Berbers, a North African people, and it is also called ''arui'' and ''waddan'' (in

Gallery

File:BarbarySheep1.jpg, Barbary sheep seeks handouts at a Texas wildlife park File:BarbarySheep2.jpg, Ewe and lamb rest in the shade of a tree File:BarbarySheep3.jpg, Lamb (closeup) File:Barbary Sheep (animal).jpg, Barbary sheep at the Wildlife Ranch in

San Antonio ("Cradle of Freedom") , image_map = , mapsize = 220px , map_caption = Interactive map of San Antonio , subdivision_type = Country , subdivision_name = United States , subdivision_type1= State , subdivision_name1 = Texas , subdivision_t ...

File:Mähnenspringer m w.jpg, Barbary sheep at Tierpark Hagenbeck, Hamburg, Germany File:Zoo003.jpg, Ewes and a juvenile at Zoologischer Garten, Berlin, Germany File:Barbary Sheep Tennoji.jpg, Barbary sheep at

Tennōji Zoo is a 11-hectare (27-acre) zoo located at Tennōji Park in Tennōji-ku, Osaka, Japan, opened on January 1, 1915. It is the third zoo to be built in Japan and is located southwest of the Shitennō-ji temple, the first Buddhist temple in Japan. ...

, Japan File:Arui02.jpg, Head of a ram File:Aoudad skeleton.jpg, Skeleton of a Barbary sheep (

Museum of Osteology The Museum of Osteology, located in Oklahoma City, Oklahoma, U.S., is a private museum devoted to the study of bones and skeletons (osteology). This museum displays over 450 skeletons of animal species from all over the world. With another 7,00 ...

) File:Barbary Sheep.png, Captive Barbary Sheep Safari West

Santa Rosa, California Santa Rosa ( Spanish for " Saint Rose") is a city and the county seat of Sonoma County, in the North Bay region of the Bay Area in California. Its estimated 2019 population was 178,127. It is the largest city in California's Wine Country and ...

References

External links

* ttps://web.archive.org/web/20051024060155/http://www.ultimateungulate.com/Artiodactyla/Ammotragus_lervia.html The Ultimate Ungulate entry on Barbary Sheep
Barbary Sheep in Texas

Barbary Sheep in Sahara
{{Taxonbar, from1=Q322141, from2=Q10732899 Caprids Mammals of Africa Mammals of North Africa Mammals of the Middle East Fauna of the Sahara Vulnerable animals Vulnerable biota of Africa Vulnerable fauna of Asia Mammals described in 1777 Taxa named by Peter Simon Pallas