Antrodemus
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''Antrodemus'' ("chamber bodied") is a dubious
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from the
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
, probably the
Morrison Formation The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandstone, siltston ...
, of Middle Park,
Colorado Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...
. It contains one species, ''Antrodemus valens'', first described and named as a species of '' Poekilopleuron'' by
Joseph Leidy Joseph Mellick Leidy (September 9, 1823 – April 30, 1891) was an American paleontologist, parasitologist and anatomist. Leidy was professor of anatomy at the University of Pennsylvania, later becoming a professor of natural history at Swarth ...
in 1870.


Discovery and species

The first described
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
specimen was a bone obtained secondhand by
Ferdinand Vandeveer Hayden Ferdinand Vandeveer Hayden (September 7, 1829 – December 22, 1887) was an American geologist noted for his pioneering surveying expeditions of the Rocky Mountains in the late 19th century. He was also a physician who served with the Union A ...
in 1869 (original discoverer unknown). It came from Middle Park, near
Granby, Colorado The Town of Granby is the List of municipalities in Colorado#Statutory town, Statutory Town that is the List of municipalities in Colorado, most populous municipality in Grand County, Colorado, Grand County, Colorado, United States. The town pop ...
, probably from Morrison Formation rocks. Hayden reported that several similar fossils had been identified as petrified
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
hooves. Hayden sent his specimen to Joseph Leidy, who identified it as half of a tail vertebra, and tentatively assigned it to the European dinosaur genus '' Poekilopleuron'' as ''Poicilopleuron'' ic''valens'', based on the shared presence of a large medullary cavity. He identified the presence of
trabeculae A trabecula (: trabeculae, from Latin for 'small beam') is a small, often microscopic, tissue element in the form of a small beam, strut or rod that supports or anchors a framework of parts within a body or organ. A trabecula generally has a ...
in ''P''. ''valens'' as a distinguishing character from ''P''. ''bucklandii'' but also noted that should better remains show more characters that could sufficiently distinguish the two taxa, it might be named ''Antrodemus''. In 1873, he amended his description and identified the species as ''Antrodemus valens''. In 1920, Charles W. Gilmore concluded that the tail vertebra named ''Antrodemus'' by Leidy was indistinguishable from those of ''
Allosaurus ''Allosaurus'' ( ) is an extinct genus of theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period ( Kimmeridgian to late Tithonian ages). The first fossil remains that could definitively be ascribed to th ...
'' and that ''Antrodemus'' should be the preferred name because, as the older name, it had priority. ''Antrodemus'' became the accepted name for this familiar genus for over fifty years until James Madsen published on the Cleveland-Lloyd specimens of ''Allosaurus'' and concluded that the ''Allosaurus'' name should be used because ''Antrodemus'' was based on material with poor, if any, diagnostic features and locality information (for example, the
geological formation A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics (lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock expo ...
that the single bone of ''Antrodemus'' came from is unknown). Subsequent authors have agreed with this assessment and have considered ''Antrodemus'' a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
''. The paleontological site that still in the present day presents the highest concentration of fossilised individuals of ''Allosaurus fragilis'' is the Cleveland-Lloyd Dinosaur Quarry, now part of the Jurassic National Monument in
Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
. The first skeletal mount obtained out of the Quarry was extracted in the late 1930s/early 1940s and finally mounted in 1961 in Guyot Hall,
Princeton University Princeton University is a private university, private Ivy League research university in Princeton, New Jersey, United States. Founded in 1746 in Elizabeth, New Jersey, Elizabeth as the College of New Jersey, Princeton is the List of Colonial ...
,
Princeton Princeton University is a private Ivy League research university in Princeton, New Jersey, United States. Founded in 1746 in Elizabeth as the College of New Jersey, Princeton is the fourth-oldest institution of higher education in the Unit ...
,
New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
, where it remained on display until 2024. For years, that skeletal mount was presented to the visitors under the taxonomic name ''Antrodemus'', before the specimens from the Cleveland-Lloyd Quarry were attributed to ''Allosaurus''.


References


External links

* * {{Taxonbar, from=Q772822 Carnosauria Dinosaur genera Kimmeridgian dinosaurs Morrison Formation Dinosaurs of the United States Fossil taxa described in 1873 Taxa named by Joseph Leidy