''Anthracotherium'', from
Ancient Greek
Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
ἄνθραξ (''ánthrax''), meaning "coal", and θηρίον (''theríon''), meaning "beast", is an
extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
artiodactyl
Artiodactyls are placental mammals belonging to the order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof). The other t ...
s characterized by having 44
teeth
A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
, with five semi-crescentic cusps on the crowns of the upper molars. The genus ranged from the middle
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
period until the early
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
, having a distribution throughout Eurasia probably even reaching South East Asia (
Kalimantan
Kalimantan (; ) is the Indonesian portion of the island of Borneo. It constitutes 73% of the island's area, and consists of the provinces of Central Kalimantan, East Kalimantan, North Kalimantan, South Kalimantan, and West Kalimantan. The non-Ind ...
and
West Timor
West Timor () is an area covering the western part of the island of Timor, except for the district of Oecussi-Ambeno (an East Timorese exclave). Administratively, West Timor is part of East Nusa Tenggara Province, Indonesia. The capital as well ...
).
Material subjectively assigned to ''Anthracotherium'' from Pakistan suggests the last species died out soon after the start of the Miocene.
Description
The genus typifies the family Anthracotheriidae, if only because it is the most thoroughly studied. In many respects, especially the anatomy of the lower jaw, ''Anthracotherium'', as with the other members of the family, is allied to the
hippopotamus
The hippopotamus (''Hippopotamus amphibius;'' ; : hippopotamuses), often shortened to hippo (: hippos), further qualified as the common hippopotamus, Nile hippopotamus and river hippopotamus, is a large semiaquatic mammal native to sub-Sahar ...
, of which it is probably an ancestral form.
The Anthracotheres, together with the hippos, are grouped with the
cetaceans
Cetacea (; , ) is an infraorder of aquatic mammals belonging to the order Artiodactyla that includes whales, dolphins and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively c ...
in the clade
Whippomorpha
Whippomorpha or Cetancodonta is a suborder of artiodactyls that contains all living cetaceans (whales, dolphins, and porpoises) and the hippopotamids. This makes it a crown group. Whippomorpha is a suborder within the order Artiodactyla (even-t ...
. Anthracotheriinae are characterized by three non-ambiguous features, which is their crown height development of the lower canine, and the presence of accessory cristulids from the hypoconulid, posthypocristulid, and labial on the lower and upper molars.
With these features into consideration, it has been found that these anthracotheres are in three dietary categories of extant herbivores: leaf browsers, fruit browsers, and grazers.
Etymology
The genus name stems from the fact that the holotype and other first specimens were originally obtained from the
Paleogene
The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...
(previously known as "
Tertiary
Tertiary (from Latin, meaning 'third' or 'of the third degree/order..') may refer to:
* Tertiary period, an obsolete geologic period spanning from 66 to 2.6 million years ago
* Tertiary (chemistry), a term describing bonding patterns in organic ch ...
")-aged
lignite
Lignite (derived from Latin ''lignum'' meaning 'wood'), often referred to as brown coal, is a soft, brown, combustible sedimentary rock formed from naturally compressed peat. It has a carbon content around 25–35% and is considered the lowest ...
beds of
Europe
Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
.
The European ''Anthracotherium magnum'' was approximately as large as a
pygmy hippo (about 2 m long and weighing up to 250
kg), but there were several smaller species and the genus also occurs in
Egypt
Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...
,
India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
and
North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
.
L'ANTHRACOTHERIUM
(in italian) Members of the genus ''Anthracotherium'', as well as other members of the family Anthracotheriidae, are known colloquially as anthracotheres.
Image:Anthracotherium magnum.jpg, ''Anthracotherium magnum''
Image:Anthracotherium coprolite.jpg, ''Anthracotherium sp.'' coprolite
A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name ...
at the Museum für Naturkunde, Berlin
References
Anthracotheriinae
Oligocene Artiodactyla
Eocene Artiodactyla
Eocene genus first appearances
Chattian genus extinctions
Eocene mammals of Asia
Eocene mammals of Europe
Extinct animals of Indonesia
Oligocene mammals of Asia
Oligocene mammals of Europe
Taxa named by Georges Cuvier
Prehistoric Artiodactyla genera
Fossil taxa described in 1822
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