Anthophora Bimaculata
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''Anthophora bimaculata'' (also called green-eyed flower bee) is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of bee.


Description

''Anthophora bimaculata'' are 8–9 mm long. The male has narrow light tergite bandages, yellow face and normally hairy middle legs clearly visible in the field. The females are Clypeus yellow, but at the base with 2 large black spots, tergite 4 and 5 gray-yellow tomentose hairs, tergias with light hair ties in the field clearly recognizable. Both genders have olive green complex eyes and one very high, clearly perceptible flight sound. Knowing the habitat of a specimen is important for identifying ''A. bimaculata''.


Range

''A. bimaculata'' have been spotted in North Africa from Morocco to Libya; a report by FRIESE (1915) from Eritrea is questionable. They also live from Portugal through southern, central and eastern Europe, Ukraine and southern Russia to eastern Siberia (Central Baikal) and across Asia Minor and the Caucasus to Iran and Kyrgyzstan; north to Central England (allegedly also recently detected in Ireland), Denmark, Latvia, Kirov; south to Sicily (also on Corsica, no evidence from Sardinia), Albania, Bulgaria (no evidence from Greece, Israel and Iran). ''A. Bimaculata'' live in central Europe in all regions in suitable habitats. Generally rare. In Germany from all federal states with the exception of Schleswig-Holstein up-to-date everywhere. Reported from all federal states in Austria with the exception of Salzburg, Tyrol and Vorarlberg. In Switzerland currently from Graubünden and Valais, historically from Lake Geneva, Lake Thun and Lake Biel as well as Ticino, Mesox and Domleschg.


Habitat

''A. bimaculata'' nests in sandy areas of lower locations: inland dunes and fields of drifting sand, sand pits, sandy embankments, ruderal areas, forest edges, and forest clearings. They nest on vegetation-free, flat surfaces or in small trailing edges. They nest only in fine and medium sands (such as drifting sand and weathered sand).


Ecology

They nest in self-dug cavities in the earth. Their nests are found both in early succession stages with only little vegetation cover as also in late stages with heavy coverage. Their nests have been found in the root balls of silver grass clusters. The main tunnel of the nest is 4–5 cm long deep in the ground. Under favorable conditions, they are found in small to large aggregations. ''A. bimaculata'' is a polylectic species, collecting pollen from 8 known plant families). Their known pollen sources are, in
Asteraceae Asteraceae () is a large family (biology), family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the Order (biology), order Asterales. The number of species in Asteraceae is rivaled only by the Orchi ...
: '' Centaurea scabiosa'', ''
Centaurea stoebe ''Centaurea stoebe'', the spotted knapweed or panicled knapweed, is a species of ''Centaurea'' native to eastern Europe, although it has spread to North America, where it is considered an invasive species. It forms a tumbleweed, helping to increa ...
'', ''
Cirsium vulgare ''Cirsium vulgare'', commonly known as spear thistle, bull thistle, or common thistle, is a species of the Asteraceae genus ''Cirsium'', native throughout most of Europe (north to 66°N, locally 68°N), Western Asia (east to the Yenisei Valley ...
, Picris hieracioides, Hypochoeris radicata''; in
Boraginaceae Boraginaceae, the Borago, borage or forget-me-not family, includes about 2,000 species of shrubs, trees, and herbs in 146 to 154 genus, genera with a worldwide distribution. The APG IV system from 2016 classifies the Boraginaceae as single famil ...
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Echium vulgare ''Echium vulgare'', known as viper's bugloss and blueweed,Dickinson, T.; Metsger, D.; Bull, J.; & Dickinson, R. (2004) ROM Field Guide to Wildflowers of Ontario. Toronto:Royal Ontario Museum, p. 203. is a species of flowering plant in the borag ...
, Anchuso officinalis''; in
Campanulaceae The family Campanulaceae (also bellflower family), of the order Asterales, contains nearly 2400 species in 84 genera of herbaceous plants, shrubs, and rarely small trees, often with milky sap. Among them are several familiar garden plants bel ...
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Trifolium arvense ''Trifolium arvense'', commonly known as the hare's-foot clover, rabbitfoot clover, stone clover or oldfield clover, is a flowering plant in the bean family Fabaceae. This species of clover is native to most of Europe, excluding the Arctic zone, ...
''; in
Hypericaceae Hypericaceae is a plant family in the order Malpighiales, comprising six to nine genera and up to 700 species, and commonly known as the St. John's wort family. Members are found throughout the world apart from extremely cold or dry habitats. ...
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Hypericum perforatum ''Hypericum perforatum'', commonly known as St. John's wort (sometimes perforate St. John's wort or common St. John's wort), is a flowering plant in the family Hypericaceae. It is a hairless, Perennial, perennial herb with woody Root, roots, y ...
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Lamiaceae The Lamiaceae ( ) or Labiatae are a family (biology), family of flowering plants commonly known as the mint, deadnettle, or sage family. Many of the plants are aromatic in all parts and include widely used culinary herbs like basil (herb), ba ...
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Teucrium scorodonia ''Teucrium scorodonia'', common name the woodland germander or wood sage, is a species of flowering plant in the genus ''Teucrium'' of the family Lamiaceae. It is native to Western Europe and Tunisia, but cultivated in many places as an ornamenta ...
''; in
Lythraceae Lythraceae is a family (biology), family of flowering plants, including 32 genus, genera, with about 620 species of Herbaceous plant, herbs, shrubs, and trees. The larger genera include ''Cuphea'' (275 spp.), ''Lagerstroemia'' (56), ''Nesaea (pl ...
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Lythrum salicaria ''Lythrum salicaria'' or purple loosestrifeFlora of NW Europe''Lythrum salicaria'' is a flowering plant belonging to the family Lythraceae. It should not be confused with other plants sharing the name loosestrife that are members of the family Pr ...
''; and in
Rosaceae Rosaceae (), the rose family, is a family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus '' Rosa''. The family includes herbs, shrubs, and trees. Most species are deciduous, but som ...
: '' Potentilla incana''. Except for the ''Hypericum'' species, all of the above serve both sexes as sources of nectar. Within the pollen loads 3 or 4 different types of pollen can be found. The field observations also showed that the flower visits on a pollen-collecting flight are randomly strung together, depending on the offer. '' Ammobates punctatus'' and '' Coelioxys rufescens'', known as
cuckoo bee The term cuckoo bee is used for a variety of different bee lineages which have evolved the kleptoparasitic behaviour of laying their eggs in the nests of other bees, reminiscent of the behavior of cuckoo birds. The name is perhaps best applied t ...
s, are parasites of ''A. bimaculata''. In England '' Coelioxys elongata'' was also observed on the nests. They are a
Univoltine Voltinism is a term used in biology to indicate the number of broods or generations of an organism in a year. The term is most often applied to insects, and is particularly in use in sericulture, where silkworm varieties vary in their voltinism. ...
species, laying one brood per year. They have a flight period from the beginning of July to the end of August, wintering as a resting larva.


Etymology

From Latin bi- "=" two "and" maculata "=" speckled ".


Taxonomy

Subgenus Heliophila KLUG, 1807. Synonyms: ''Saropoda rotundata'' (PANZER, 1798); ''Heliophila bimaculata'' (PANZER, 1798).


References

{{Taxonbar, from=Q292895 Apinae Insects described in 1798