Anthocorid
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Anthocoridae is a family of bugs, commonly called minute pirate bugs or flower bugs. Worldwide there are 500-600 species.


Description

Anthocoridae are 1.5–5 mm long and have soft, elongated oval, flat bodies, often patterned in black and white. The head is extended forward and the antennae are longer than the head and visible from above. They possess a piercing and sucking three-segmented beak or labium used to inject prey with digestive enzymes and consume food. In general appearance, they resemble common plant bugs (
Miridae The Miridae are a large and diverse insect family at one time known by the Synonym (taxonomy), taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or "mirid bugs". Common names include plant bugs, leaf bugs, and g ...
), but Anthocoridae differ by their possession of two ocelli as adults. Anthocorids possess two pairs of wings with hemelytra and membranous hindwings. Many species are referred to as insidious flower bugs or pirate bugs. The scientific name is a combination of the Greek words ''anthos'' "flower" and ''koris'' "bug".


Habitat and behaviour

Many species can be found in cryptic habitats such as
galls Galls (from the Latin , 'oak-apple') or ''cecidia'' (from the Greek , anything gushing out) are a kind of swelling growth on the external tissues of plants. Plant galls are abnormal outgrowths of plant tissues, similar to benign tumors or wart ...
, but can also be present in open surface environments. They can often be found in many agricultural crops. They can feed on plant material, but mostly feed on other small soft-bodied
arthropods Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
. Anthocorids are often predacious both as nymphs and adults. They are beneficial as
biological control Biological control or biocontrol is a method of controlling pests, whether pest animals such as insects and mites, weeds, or pathogens affecting animals or plants by using other organisms. It relies on predation, parasitism, herbivory, or o ...
agents. ''
Orius insidiosus ''Orius insidiosus'', common name the insidious flower bug, is a species of Anthocoridae, minute pirate bug, a predatory insect in the order (biology), order Hemiptera (the true bugs). They are considered beneficial, as they feed on small pest art ...
'', the "insidious flower bug", for example, feeds on the eggs of the corn earworm (''Helicoverpa zea''). ''Orius insidiosus'' is often released in greenhouses against mites and
thrips Thrips (Order (biology) , order Thysanoptera) are minute (mostly long or less), slender insects with fringed wings and unique asymmetrical mouthparts. Entomologists have species description , described approximately 7,700 species. They fly on ...
. Eggs are laid in plant material and hatch in approximately 3 to 5 days. Nymphs require at least 20 days to progress through five
instars An instar (, from the Latin ''wikt:instar#Latin, īnstar'' 'form, likeness') is a developmental stage of arthropods, such as insects, which occurs between each ecdysis, moult (''ecdysis'') until sexual maturity is reached. Arthropods must shed the ...
. Adults live for approximately 35 days. These small insects can bite humans, however, they do not feed on human blood or inject venom or saliva. Reactions to bites in individuals can range from no effect to minor swelling and irritation.


Systematics

There are two subfamilies and at least 8 tribes:


Anthocorinae

Auth. Fieber, 1837 ;tribe Almeidini Carayon, 1972 # '' Almeida'' Distant, 1910 # '' Australmeida'' Woodward, 1977 # ''
Lippomanus ''Lippomanus''Distant WL (1904) ''Ann. Mag. Nat. Hist.'', (7) 14. is a genus of seed bugs in the subfamily Anthocorinae (tribe Almeidini), erected by William Lucas Distant William Lucas Distant (12 November 1845 Rotherhithe – 4 February ...
'' Distant, 1904 ;tribe
Anthocorini Anthocorini is a tribe of minute pirate bugs in the family Anthocoridae. There are more than 30 described species in Anthocorini. Genera ''BioLib'' includes the following in tribe Anthocorini: # ''Acompocoris'' Reuter, 1875 # ''Anthocoris'' Fal ...
Fieber, 1837 Selected genera: * ''
Acompocoris ''Acompocoris'' is a genus of minute pirate bugs in the family Anthocoridae Anthocoridae is a family of bugs, commonly called minute pirate bugs or flower bugs. Worldwide there are 500-600 species. Description Anthocoridae are 1.5–5  ...
'' Reuter, 1875 i c g b * ''
Anthocoris ''Anthocoris'' is a genus of minute pirate bugs in the family Anthocoridae. There are at least 30 described species in ''Anthocoris''. Species These 32 species belong to the genus ''Anthocoris'': * '' Anthocoris albiger'' Reuter, 1884 * '' An ...
'' Fallen, 1814 i c g b * '' Coccivora'' McAtee & Malloch, 1925 i c g b * ''
Dufouriellus ''Dufouriellus'' is a monotypic genus of minute pirate bugs in the tribe Dufouriellini. The described species is ''Dufouriellus ater'', which has been recorded from much of western Europe through to Bohemia, Moravia, Slovakia, southern Scandinavi ...
'' Kirkaldy, 1906 * ''
Elatophilus ''Elatophilus'' is a genus of minute pirate bugs in the family Anthocoridae Anthocoridae is a family of bugs, commonly called minute pirate bugs or flower bugs. Worldwide there are 500-600 species. Description Anthocoridae are 1.5–5  ...
'' Reuter, 1884 i c g b * ''
Melanocoris ''Melanocoris'' is a genus of minute pirate bugs in the family Anthocoridae Anthocoridae is a family of bugs, commonly called minute pirate bugs or flower bugs. Worldwide there are 500-600 species. Description Anthocoridae are 1.5–5  ...
'' Champion, 1900 i c g b * ''
Temnostethus ''Temnostethus'' is a genus of minute pirate bugs in the family Anthocoridae. There are about eight described species in ''Temnostethus''. Species These eight species belong to the genus ''Temnostethus'': * ''Temnostethus dacicus'' (Puton, 1888) ...
'' Fieber, 1860 i c g b * ''
Tetraphleps ''Tetraphleps'' is a genus of minute pirate bugs in the family Anthocoridae. There are about 10 described species in ''Tetraphleps''. Species These 10 species belong to the genus ''Tetraphleps'': * ''Tetraphleps aterrima'' (J.Sahlberg, 1878) * ...
'' Fieber, 1860 i c g b ;tribe Blaptopstethini Carayon, 1972 # '' Blaptostethoides'' Carayon, 1972 # '' Blaptostethus'' Fieber, 1860 ;tribe
Dufouriellini Dufouriellini is a tribe of minute pirate bugs in the family Anthocoridae. There are about 7 genera and 15 described species in Dufouriellini. Genera * '' Alofa'' Herring, 1976 * '' Amphiareus'' Distant, 1904 * '' Brachysteles'' Mulsant & Rey, ...
Van Duzee, 1916 Selected genera: * ''
Amphiareus ''Amphiareus'' is a genus of minute pirate bugs belonging to the family Anthocoridae Anthocoridae is a family of bugs, commonly called minute pirate bugs or flower bugs. Worldwide there are 500-600 species. Description Anthocoridae are 1.5†...
'' Distant, 1904 * ''
Brachysteles ''Brachysteles'' is a genus of minute pirate bugs in the family Anthocoridae. There are at least three described species in ''Brachysteles''. Species These three species belong to the genus ''Brachysteles'': * '' Brachysteles espagnoli'' Ribes, ...
'' Mulsant & Rey, 1852 * ''
Buchananiella ''Buchananiella'' is a genus of minute pirate bugs in the family Anthocoridae. There are at least three described species in ''Buchananiella''. Species These three species belong to the genus ''Buchananiella'': * '' Buchananiella continua'' (Wh ...
'' Reuter, 1884 * ''
Cardiastethus ''Cardiastethus'' is a genus of minute pirate bugs in the family Anthocoridae. There are more than 20 described species in ''Cardiastethus''. Species These 23 species belong to the genus ''Cardiastethus'': * '' Cardiastethus assimilis'' (Reuter ...
'' Fieber, 1860 ;tribe
Oriini Oriini is a tribe of minute pirate bugs in the family Anthocoridae. There are about 6 genera and more than 60 described species in Oriini. Genera These six genera belong to the tribe Oriini: * '' Macrothacheliella'' Champion, 1900 * '' Macrotra ...
Carayon, 1955 Selected genera: * ''
Orius The genus ''Orius'' (commonly called minute pirate bug) consists of omnivorous bugs in the family Anthocoridae (pirate bugs). Adults are 2–5 mm long and feed mostly on smaller insects, larva and eggs, such as spider mites, thrips, ju ...
'' Wolff, 1811 i c g b ;tribe Scolopini Carayon, 1954 Selected genera: * ''
Calliodis ''Calliodis'' is a genus of bugs in the family Anthocoridae Anthocoridae is a family of bugs, commonly called minute pirate bugs or flower bugs. Worldwide there are 500-600 species. Description Anthocoridae are 1.5–5 mm long and have ...
'' Reuter, 1871 * '' Scoloposcelis'' Fieber, 1864 ;tribe Xylocorini Carayon, 1972 (monotypic) # '' Xylocoris'' Dufour, 1831 ;''incertae sedis'' # '' Cyrtosternum'' Fieber, 1860


Lasiochilinae

Auth. Carayon, 1972 (sometimes placed at family level: "Lasiochilidae") * tribe Lasiochilini Carayon, 1972 # ''
Lasiochilus ''Lasiochilus'' is a genus of true bugs in the subfamily Lasiochilinae and tribe Lasiochilini. There are about 17 described species in ''Lasiochilus''. Species These 17 species belong to the genus ''Lasiochilus'': * ''Lasiochilus ather'' Herrin ...
'' Reuter, 1871 # '' Plochiocoris'' Champion, 1900 * genera ''incertae sedis''


Unplaced genera

# ''
Lilia Lilia (Latin plural, meaning "lilies" in English; singular, ''lilium'') are pit traps arranged in a quincunx pattern dug by the Roman armies in front of their defences. Frequently they had sharpened stakes set inside them as an extra obstacle to ...
'' White, 1879 # '' Opisthypselus'' Reuter, 1908 * 4 fossil genera Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.netPéricart, J. 1996. Family Anthocoridae Fieber, 1836 flower bugs, minute pirate bugs, pp. 108–318. In Aukema, B. and C. Rieger, eds. ''Catalogue of the Heteroptera of the Palaearctic Region''. Vol. 2. Cimicomorpha I. Netherlands Entomological Society, Amsterdam. 359 pp.


References


External links


Pirate bugsPirate bugs
(with pictures) * {{Authority control Heteroptera families