''Antarctosuchus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of capitosauroid

temnospondyl Temnospondyli (from Greek language, Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') or temnospondyls is a diverse ancient order (biology), order of small to giant tetrapods—often considered Labyrinth ...

known from the

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epoch (geology), epochs of the Triassic period (geology), period or the middle of three series (stratigraphy), series in which the Triassic system (stratigraphy), system is di ...

upper Fremouw Formation in the

Transantarctic Mountains The Transantarctic Mountains (abbreviated TAM) comprise a mountain range of uplifted rock (primarily sedimentary) in Antarctica which extends, with some interruptions, across the continent from Cape Adare in northern Victoria Land to Coats L ...

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...

. It contains a single species, ''Antarctosuchus polyodon''.

Discovery

''Antarctosuchus'' was named by Christian A. Sidor, J. Sébastien Steyer and William R. Hammer in

2014 The year 2014 was marked by the surge of the Western African Ebola epidemic, West African Ebola epidemic, which began in 2013, becoming the List of Ebola outbreaks, most widespread outbreak of the Ebola, Ebola virus in human history, resul ...

and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Antarctosuchus polyodon''. The generic name refers to

, where the only known specimen was collected, and ''suchus'', a common suffix for

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

ized from the

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

''souchos'', an Egyptian crocodile god. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''polyodon'', is derived from the Greek πολύς (''polús'') "much", "many" and ὀδών (''odon''), "tooth", in reference to its unique

teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

morphology. ''Antarctosuchus'' is known solely from its

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

, a large and relatively complete

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

. It was collected form the upper Fremouw Formation in the central

, dating to the

. To date, ''Antarctosuchus polyodon'' is the only other

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

species from the formation apart from '' Kryostega collinsoni'', although indeterminate remains referred to benthosuchids and a cranial fragment assigned to ''

Parotosuchus ''Parotosuchus'' is an extinct genus of capitosaurian temnospondyls within the family Mastodonsauridae. Fossils are known from the Early Triassic of Europe, Africa, Australia, and Antarctica. It was about long and likely lived in aquatic envir ...

sp.'' have also been collected.

Description

As its specific name suggests, ''Antarctosuchus polyodon'' is distinguished by the possession of numerous, extremely small

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

ry,

palatine A palatine or palatinus (Latin; : ''palatini''; cf. derivative spellings below) is a high-level official attached to imperial or royal courts in Europe since Roman Empire, Roman times.

, and ectopterygoid teeth, a dental pattern that suggests specialization on small prey, possibly

invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...

s. Other autapomorphic features of this taxon are a parachoanal tooth row that extends far posterior to the choana and occipital condyles which are set close to the midline. ''Antarctosuchus'' possesses a relatively flat skull, low

occiput The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone lies over the occipital lobes of the ...

and well-developed sensory canals, suggesting an aquatic lifestyle. A

phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...

places ''Antarctosuchus'' within

Capitosauroidea The Mastodonsauroidea are an extinct superfamily of temnospondyl amphibians known from the Triassic. Fossils belonging to this superfamily have been found in North America, Greenland, Europe, Asia, and Australia. '' Ferganobatrachus'' from the ...

, a clade of derived Triassic stereospondyls as the

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to '' Paracyclotosaurus crookshanki'' from the Triassic Denwai Formation of

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

References

{{Taxonbar, from=Q16961664 Capitosauria Triassic temnospondyls of Antarctica Fossil taxa described in 2014