The Anserinae are a

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

in the

waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which i ...

family

Anatidae The Anatidae are the biological family (biology), family of water birds that includes ducks, goose, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted f ...

. It includes the

swan Swans are birds of the genus ''Cygnus'' within the family Anatidae. The swans' closest relatives include the goose, geese and ducks. Swans are grouped with the closely related geese in the subfamily Anserinae where they form the tribe (biology) ...

s and the true

geese A goose (: geese) is a bird of any of several waterfowl species in the family Anatidae. This group comprises the genera '' Anser'' (grey geese and white geese) and ''Branta'' (black geese). Some members of the Tadorninae subfamily (e.g., Egyp ...

. Under alternative systematical concepts (see e.g., Terres & NAS, 1991), it is split into two subfamilies, the Anserinae contain the geese and the ducks, while the Cygninae contain the swans.

Systematics

Swan Swans are birds of the genus ''Cygnus'' within the family Anatidae. The swans' closest relatives include the goose, geese and ducks. Swans are grouped with the closely related geese in the subfamily Anserinae where they form the tribe (biology) ...

s (Tribe

Cygnini Swans are birds of the genus ''Cygnus'' within the family Anatidae. The swans' closest relatives include the geese and ducks. Swans are grouped with the closely related geese in the subfamily Anserinae where they form the tribe Cygnini. Sometim ...

) *Genus '' Cygnus'' – true swans: The black-and-yellow-billed swans are sometimes separated in the genus ''Olor''. *Genus †''

Afrocygnus ''Afrocygnus'' is an extinct genus of swan, which lived during the Late Miocene, and perhaps up to the Late Pliocene, in what is today North Africa. The only genus of swan known in Africa, aside from fragmentary Pleistocene remains found in East ...

'' (

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

North Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...

) *Genus †''

Annakacygna ''Annakacygna'' is a genus of flightless marine swan from the Miocene of Japan. Named in 2022, ''Annakacygna'' displays a series of unique adaptations setting it apart from any other known swan, including a filter feeding lifestyle, a highly mobi ...

–'' short-winged swans (Miocene of

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

) *Genus †''

Megalodytes ''Megalodytes'' is an extinct genus of waterfowl. The only known species, ''M. morejohni'', is known from fragmentary wingbones found in Middle Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and e ...

'' (Miocene of

California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...

) True geese (Tribe Anserini) *Genus '' Anser'' – grey and white geese *Genus ''

Branta The black geese of the genus ''Branta'' are waterfowl belonging to the Goose, true geese and swans subfamily Anserinae. They occur in the northern coastal regions of the Palearctic and all over North America, Bird migration, migrating to more so ...

'' – black geese (including †'' B. rhuax'', formerly placed in ''Geochen'') Unresolved *Genus ''

Coscoroba The coscoroba swan (''Coscoroba coscoroba'') is a species of waterfowl in the subfamily Anserinae of the family Anatidae.HBW and BirdLife International (2021) Handbook of the Birds of the World and BirdLife International digital checklist of the ...

'' – coscoroba swan These two genera are distinct from other geese and often elevated to a subfamily of their own (Cereopsinae), or alternatively into the

shelduck The shelducks, most species of which are found in the genus ''Tadorna'' (except for the Radjah shelduck, which is now found in its own monotypic genus ''Radjah''), are a group of large birds in the Tadorninae subfamily of the Anatidae, the biolog ...

subfamily Tadorninae: Tribe Cereopseini *Genus ''

Cereopsis The Cape Barren goose (''Cereopsis novaehollandiae''), sometimes also known as the pig goose, is a species of goose endemic to southern Australia. It is a distinctive large, grey bird that is mostly terrestrial and is not closely related to oth ...

'' – Cape Barren goose *Genus †''

Cnemiornis The New Zealand goose is a bird of the extinct genus ''Cnemiornis'' of the family Anatidae, subfamily Anserinae. The genus, endemic to New Zealand, consisted of two species: the North Island goose, ''C. gracilis'' and the South Island goose ''C. ...

'' – New Zealand geese (

prehistoric Prehistory, also called pre-literary history, is the period of human history between the first known use of stone tools by hominins million years ago and the beginning of recorded history with the invention of writing systems. The use o ...

) Some enigmatic

subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s of very large goose-like birds from the

Hawaiian Islands The Hawaiian Islands () are an archipelago of eight major volcanic islands, several atolls, and numerous smaller islets in the Pacific Ocean, North Pacific Ocean, extending some from the Hawaii (island), island of Hawaii in the south to nort ...

do not appear to be

moa-nalo The moa-nalo are a group of extinct aberrant, goose-like ducks that lived on the larger Hawaiian Islands, except Hawaii (island), Hawaii itself, in the Pacific. They were the major herbivores on most of these islands until they became extinct a ...

s (goose-sized

dabbling duck The Anatinae are a subfamily of the family Anatidae (swans, geese and ducks). Its surviving members are the dabbling ducks, which feed mainly at the surface rather than by diving. The other members of the Anatinae are the extinct moa-nalo, a y ...

s) or ''B. rhuax''. They cannot be assigned to any genus living today, though both may be fairly close to ''Branta'': * Very large Hawaiʻian goose, ?''Branta'' sp. * Giant Oʻahu goose,

sp. et gen. indet.

References

* Terres, John K. &

National Audubon Society The National Audubon Society (Audubon; ) is an American non-profit environmental organization dedicated to conservation of birds and their habitats. Located in the United States and incorporated in 1905, Audubon is one of the oldest of such orga ...

(1991): ''The Audubon Society Encyclopedia of North American Birds''. Wings Books, New York.

Systematics

References

Further reading