''Anningia'' is an extinct genus in

Varanopidae Varanopidae is an extinct family (biology), family of amniotes known from the Late Carboniferous to Middle Permian that resembled monitor lizards (with the name of the group deriving from the monitor lizard genus ''Varanus'') and may have filled ...

, a family of

monitor lizard Monitor lizards are lizards in the genus ''Varanus,'' the only extant genus in the family Varanidae. They are native to Africa, Asia, and Oceania, and West African Nile monitor, one species is also found in south America as an invasive species. A ...

-like

amniotes Amniotes are tetrapod vertebrate animals belonging to the clade Amniota, a large group that comprises the vast majority of living terrestrial and semiaquatic vertebrates. Amniotes evolved from amphibious stem tetrapod ancestors during the ...

. It contains a single species, ''Anningia megalops''. The species was first described by

Robert Broom Robert Broom Fellow of the Royal Society, FRS FRSE (30 November 1866 6 April 1951) was a British- South African medical doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University ...

in 1927 as a transitional species between early

synapsid Synapsida is a diverse group of tetrapod vertebrates that includes all mammals and their extinct relatives. It is one of the two major clades of the group Amniota, the other being the more diverse group Sauropsida (which includes all extant rept ...

s and later occurring therapsids. While it was later suggested that the

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

material, a partial poorly preserved

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

, was not diagnosable and that the species name is ''

nomen vanum This is a list of terms and symbols used in scientific names for organisms, and in describing the names. For proper parts of the names themselves, see List of Latin and Greek words commonly used in systematic names. Many of the abbreviations are ...

''. It was assigned to Varanopidae (subfamily Mesenosaurinae) in 2018. The fossil is from the

Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the s ...

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...

References

Prehistoric synapsid genera Fossil taxa described in 1927 {{tetrapod-stub