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''Angophora'' is a genus of nine species of trees and shrubs in the myrtle family,
Myrtaceae Myrtaceae (), the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All ...
.
Endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to eastern Australia, they differ from other
eucalypt Eucalypt is any woody plant with Capsule (fruit), capsule fruiting bodies belonging to one of seven closely related genera (of the tribe Eucalypteae) found across Australia: ''Eucalyptus'', ''Corymbia'', ''Angophora'', ''Stockwellia'', ''Allosyn ...
s in having juvenile and adult leaves arranged in opposite pairs,
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s reduced to projections on the edge of the floral cup, four or five overlapping, more or less round
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s, and a papery or thin, woody, often strongly ribbed capsule. Species are found between the Atherton Tableland in Queensland and south through New South Wales to eastern Victoria, Australia.


Description

Plants in the genus ''Angophora'' are trees, occasionally shrubs, with rough bark except for '' A. costata''. The juvenile leaves differ from adult leaves in being hairy with raised oil
glands A gland is a Cell (biology), cell or an Organ (biology), organ in an animal's body that produces and secretes different substances that the organism needs, either into the bloodstream or into a body cavity or outer surface. A gland may also funct ...
. Both juvenile and adult leaves are arranged in opposite pairs, the adult leaves usually glabrous and paler on the lower surface. The flower buds are arranged in groups of three or seven. The flower has four or five small sepals, reduced to small projections on the rim of the floral cup. There are four or five more or less round, keeled, overlapping petals and whorls of many creamy white
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s. Unlike in ''
Eucalyptus ''Eucalyptus'' () is a genus of more than 700 species of flowering plants in the family Myrtaceae. Most species of ''Eucalyptus'' are trees, often Mallee (habit), mallees, and a few are shrubs. Along with several other genera in the tribe Eucalyp ...
'' and ''
Corymbia ''Corymbia'', commonly known as bloodwoods, is a genus of about one hundred species of tree that, along with ''Eucalyptus'', '' Angophora'' and several smaller groups, are referred to as eucalypts. Until 1990, corymbias were included in the g ...
'', the petals and sepals are not fused to form a cap-like operculum. The fruit is a papery or thin, slightly woody, hairy capsule with longitudinal ribs.


Taxonomy

The genus ''Angophora'' was first formally described in 1797 by Antonio José Cavanilles in his book ''Icones et Descriptiones Plantarum''. ''Angophora'' is closely related to ''
Corymbia ''Corymbia'', commonly known as bloodwoods, is a genus of about one hundred species of tree that, along with ''Eucalyptus'', '' Angophora'' and several smaller groups, are referred to as eucalypts. Until 1990, corymbias were included in the g ...
'' and ''
Eucalyptus ''Eucalyptus'' () is a genus of more than 700 species of flowering plants in the family Myrtaceae. Most species of ''Eucalyptus'' are trees, often Mallee (habit), mallees, and a few are shrubs. Along with several other genera in the tribe Eucalyp ...
'', and all three genera are often referred to as "eucalypts". Collectively the eucalypts dominate many Australian
ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...
s. Taxonomists have long recognised the relationships between the eucalypt taxa, but have not agreed upon a classification scheme. Some have proposed merging ''Angophora'' and ''Corymbia'' into genus ''Eucalyptus'' as
subgenera In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
, a plan which was immediately rejected by others. Some authors maintain ''Angophora'' as a genus, while others continue to debate the issue.Nelson, G., et al. (2003)
Brummitt on paraphyly: a response.
''Taxon'' 52(2), 295-98.
Among the eucalypts, ''Angophora'' species were nicknamed "apples" by European settlers, who thought they resembled apple trees. Many are still known commonly as apples today.


Species

The following is a list species of accepted by the
Australian Plant Census The Australian Plant Census (APC) provides an online interface to currently accepted, published, scientific names of the vascular flora of Australia, as one of the output interfaces of the national government Integrated Biodiversity Information Sys ...
as at March 2020: * '' Angophora bakeri'' E.C.Hall - NSW ** ''Angophora bakeri'' E.C.Hall subsp. ''bakeri'' - NSW ** ''Angophora bakeri'' subsp. ''crassifolia'' G.J.Leach - NSW * '' Angophora costata'' ( Gaertn.) Britten – Qld, NSW ** ''Angophora costata'' (Gaertn.) Britten subsp. ''costata'' – Qld, NSW ** ''Angophora costata'' subsp. ''euryphylla'' L.A.S.Johnson ex G.J.Leach – NSW * '' Angophora floribunda'' ( Sm.) Sweet - Qld, NSW, Vic. * '' Angophora hispida'' (Sm.) Blaxell - NSW * '' Angophora inopina'' K.D.Hill - NSW * ''
Angophora leiocarpa ''Angophora leiocarpa'', commonly known as rusty gum, is a species of small to medium-sized tree that is endemic to eastern Australia. It has smooth bark on the trunk and branches, lance-shaped adult leaves, flower buds usually in groups of thre ...
'' (L.A.S.Johnson ex G.J.Leach) K.R.Thiele & Ladiges - NSW, Qld. * '' Angophora melanoxylon'' R.T.Baker - Qld., NSW * '' Angophora robur'' L.A.SJohnson & K.D.Hill - NSW * '' Angophora subvelutina'' F.Muell. - NSW * '' Angophora woodsiana'' F.M.Bailey - Qld., NSW


Distribution

Angophoras are found in coastal Queensland, New South Wales and Victoria from the Atherton Tableland to eastern Victoria. Angophora hispida DSC 7157 (11023015645).jpg , '' A. hispida'' buds & flowers Angophora hispida DSC 6592 (33368522242).jpg, '' A. hispida'' fruit Angophora hispida Angophora hispida new growth IMG 5611 (1867066136).jpg, '' A. hispida'' lignotuber regrowth after fire Angophora hispida DSC 4503 (27666550975).jpg, '' A. hispida'' opposite and decussate leaves


References


External links

{{Taxonbar, from=Q1904772 Endemic flora of Australia Myrtaceae genera Taxa named by Antonio José Cavanilles Myrtales of Australia