Amoebidium
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''Amoebidium'' is a genus of
unicellular A unicellular organism, also known as a single-celled organism, is an organism that consists of a single cell, unlike a multicellular organism that consists of multiple cells. Organisms fall into two general categories: prokaryotic organisms and ...
,
symbiotic Symbiosis (Ancient Greek : living with, companionship < : together; and ''bíōsis'': living) is any type of a close and long-term biolo ...
eukaryote The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms ...
s in the
Opisthokont The opisthokonts () are a broad group of eukaryotes, including both the animal and fungus kingdoms. The opisthokonts, previously called the "Fungi/Metazoa group", are generally recognized as a clade. Opisthokonts together with Apusomonadida and ...
group
Mesomycetozoea The Ichthyosporea (or DRIP clade, or Mesomycetozoea) are a small group of Opisthokonta in Eukaryota (formerly protists), mostly parasites of fish and other animals. Significance They are not particularly distinctive morphologically, appearing i ...
, family Amoebidiidae.Reynolds, N.K., M.E. Smith, E.D. Tretter, J. Gause, D. Heeney, M.J. Cafaro, J.F. Smith, S.J. Novak, W.A. Bourland, M.M. White. 2017. Resolving relationships at the animal-fungal divergence: A molecular phylogenetic study of the protist trichomycetes (Ichthyosporea, Eccrinida). Molecular Phylogenetics and Evolution 109, 447-464. ''Amoebidium'' species attach to the
exoskeleton An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the internal organs, in contrast to an internal endoskeleton (e.g. human skeleton, that ...
of freshwater aquatic
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s such as midge larvae and water fleas (''
Daphnia ''Daphnia'' is a genus of small planktonic crustaceans, in length. ''Daphnia'' are members of the Order (biology), order Anomopoda, and are one of the several small aquatic crustaceans commonly called water fleas because their Saltation (gait), ...
'').Lichtwardt, R.W., M.J. Cafaro, M.M. White. 2001. The Trichomycetes: Fungal Associates of Arthropods Revised Edition. Published online http://www.nhm.ku.edu/%7Efungi/Monograph/Text/Mono.htm The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is
Amoebidium parasiticum
', which is also one of the only species to be cultured
axenic In biology, axenic (, ) describes the state of a culture in which only a single species, variety, or strain of organism is present and entirely free of all other contaminating organisms. The earliest axenic cultures were of bacteria or unicellul ...
ally.Whisler, H.C., 1960. Pure culture of the Trichomycete, Amoebidium parasiticum. Nature 186, 732-733.


Etymology

Derived from the word "
amoeba An amoeba (; less commonly spelled ameba or amœba; : amoebas (less commonly, amebas) or amoebae (amebae) ), often called an amoeboid, is a type of Cell (biology), cell or unicellular organism with the ability to alter its shape, primarily by ...
" to refer to the amoeba-like dispersal cells formed during some stages of the life cycle.


Description

''Amoebidium'' species are single-celled,
cigar A cigar is a rolled bundle of dried and Fermentation, fermented tobacco leaves made to be Tobacco smoking, smoked. Cigars are produced in a variety of sizes and shapes. Since the 20th century, almost all cigars are made of three distinct comp ...
-shaped or tubular in vegetative growth form (=
thallus Thallus (: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. A thallus usually names the entir ...
), and attach to the
exoskeleton An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the internal organs, in contrast to an internal endoskeleton (e.g. human skeleton, that ...
of various freshwater arthropod hosts ( Crustaecea or
Insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
a) by means of a secreted, glue-like basal
holdfast Holdfast most often refers to: *Holdfast (biology), a root-like structure that anchors aquatic sessile organisms to their substrate *Holdfast (tool), a tool used to secure a workpiece to a workbench or anvil Holdfast or hold fast may also refer t ...
. The thalli are coenocytic (i.e. lack divisions within the cell) and are unbranched.
Sexual reproduction Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote tha ...
is unknown.
Asexual reproduction Asexual reproduction is a type of reproduction that does not involve the fusion of gametes or change in the number of chromosomes. The offspring that arise by asexual reproduction from either unicellular or multicellular organisms inherit the f ...
may proceed along two different routes: 1) the entire content of the cell divides into elongated,
uninucleate {{Short pages monitor