Amalda Acutapex
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''Amalda'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of medium-sized
sea snail Sea snails are slow-moving marine (ocean), marine gastropod Mollusca, molluscs, usually with visible external shells, such as whelk or abalone. They share the Taxonomic classification, taxonomic class Gastropoda with slugs, which are distinguishe ...
s, a marine
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
in the family
Ancillariidae The Ancillariidae is a taxonomic family of sea snails, marine gastropod mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, ...
, the olives and allies.Bouchet, P. (2014). Amalda H. Adams & A. Adams, 1853. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=390993 on 2015-01-18


Description

The thin shell is not umbilicated. The
spire A spire is a tall, slender, pointed structure on top of a roof of a building or tower, especially at the summit of church steeples. A spire may have a square, circular, or polygonal plan, with a roughly conical or pyramidal shape. Spire ...
is elongated with the suture enamelled. The primary spire callus covers most of the spire, including at least part of the protoconch, leaving in most cases only the first protoconch whorl exposed. The inner
lip The lips are a horizontal pair of soft appendages attached to the jaws and are the most visible part of the mouth of many animals, including humans. Mammal lips are soft, movable and serve to facilitate the ingestion of food (e.g. sucklin ...
has the callus moderate, defined. The outer lip is simple.


Distribution

These snails usually live in the sand in fairly shallow water in tropical and temperate regions of the world with a particularly rich fauna in Australia and the Indo-West Pacific. Most species are predators of marine
bivalves Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed by a calcified exoskeleton consis ...
. In general, both shallow and deep-water species of ''Amalda'' tend to have relatively narrow distribution ranges within the Indo-Pacific, where local faunas are often dominated by narrow endemics, particularly in South Africa and, to a lesser extent, Australia. While some species exhibit broader distributions—such as ''A. hinomotoensis'', found from Japan to Taiwan, and ''A. mamillata'', present in regions including
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
, the
Strait of Malacca The Strait of Malacca is a narrow stretch of water, long and from wide, between the Malay Peninsula to the northeast and the Indonesian island of Sumatra to the southwest, connecting the Andaman Sea (Indian Ocean) and the South China Sea (Pa ...
,
Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
, and the
China Seas The China Seas consist of a series of marginal seas in the Western Pacific Ocean, around China. They are the major components signifying the transition from the continent of Asia to the Pacific Ocean.Pinxian Wang, Qianyu Li, Chun-Feng Li, ''Geol ...
(Gratecap 2016b)—such broad distributions should be interpreted with caution.Yuri I. Kantor, Magalie Castelin, Alexander Fedosov, Philippe Bouchet (2020), ''The Indo-Pacific Amalda (Neogastropoda, Olivoidea, Ancillariidae) revisited with molecular data, with special emphasis on New Caledonia''; European Journal of Taxonomy, [S.l.], n. 706, aug. 2020. ISSN 2118-9773


Species

Species within the genus ''Amalda'' include: * † ''
Amalda abessensis ''Amalda'' is a genus of medium-sized sea snails, a marine gastropod mollusc in the family Ancillariidae, the olives and allies.Bouchet, P. (2014). Amalda H. Adams & A. Adams, 1853. Accessed through: World Register of Marine Species at http://ww ...
'' Lozouet, 1992 * '' Amalda abyssicola'' Schepman, 1911 * '' Amalda acuta'' Ninomiya, 1991 * † '' Amalda acutapex'' Raven & Recourt, 2018 * † ''
Amalda adelaidensis ''Amalda'' is a genus of medium-sized sea snails, a marine gastropod mollusc in the family Ancillariidae, the olives and allies.Bouchet, P. (2014). Amalda H. Adams & A. Adams, 1853. Accessed through: World Register of Marine Species at http://ww ...
'' (Ludbrook, 1958) * '' Amalda alabaster'' Kantor, Castelin, Fedosov & Bouchet, 2020 * '' Amalda albanyensis'' Ninomiya, 1987 * '' Amalda albocallosa'' (Lischke, 1873) * † ''
Amalda allani ''Amalda'' is a genus of medium-sized sea snails, a marine gastropod mollusc in the family Ancillariidae, the olives and allies.Bouchet, P. (2014). Amalda H. Adams & A. Adams, 1853. Accessed through: World Register of Marine Species at http://ww ...
'' (Olson, 1956) * '' Amalda allaryi'' Bozetti, 2007 * '' Amalda angustata'' (G.B. Sowerby II, 1859) * † ''
Amalda anomala ''Amalda'' is a genus of medium-sized sea snails, a marine gastropod mollusc in the family Ancillariidae, the olives and allies.Bouchet, P. (2014). Amalda H. Adams & A. Adams, 1853. Accessed through: World Register of Marine Species at http://ww ...
'' (Schlotheim, 1820) * † ''
Amalda arenaria ''Amalda'' is a genus of medium-sized sea snails, a marine gastropod mollusc in the family Ancillariidae, the olives and allies.Bouchet, P. (2014). Amalda H. Adams & A. Adams, 1853. Accessed through: World Register of Marine Species at http://ww ...
'' (Cossmann, 1889) * † ''
Amalda aturensis ''Amalda'' is a genus of medium-sized sea snails, a marine gastropod mollusc in the family Ancillariidae, the olives and allies.Bouchet, P. (2014). Amalda H. Adams & A. Adams, 1853. Accessed through: World Register of Marine Species at http://ww ...
'' (Peyrot, 1928) * † ''
Amalda auberi ''Amalda'' is a genus of medium-sized sea snails, a marine gastropod mollusc in the family Ancillariidae, the olives and allies.Bouchet, P. (2014). Amalda H. Adams & A. Adams, 1853. Accessed through: World Register of Marine Species at http://ww ...
'' Chavan, 1965 * '' Amalda aureocallosa'' Shikama & Oishi, 1977 * '' Amalda aureomarginata'' Kilburn & Bouchet, 1988 * '' Amalda aureus'' Ninomiya, 1990 * ''
Amalda australis ''Amalda australis'', common name the southern olive, is a medium-sized sea snail, a gastropod mollusc of the family (biology), family Ancillariidae.Sartori, A.; Rosenberg, G. (2014). Amalda australis (G.B. Sowerby I, 1830). Accessed through: Wor ...
'' (G.B. Sowerby I, 1830) * '' Amalda bathamae'' (
Dell Dell Inc. is an American technology company that develops, sells, repairs, and supports personal computers (PCs), Server (computing), servers, data storage devices, network switches, software, computer peripherals including printers and webcam ...
, 1956)
* '' Amalda beachportensis'' (Verco, 1909) * '' Amalda bellonarum'' Kilburn & Bouchet, 1988 * '' Amalda benthicola'' (
Dell Dell Inc. is an American technology company that develops, sells, repairs, and supports personal computers (PCs), Server (computing), servers, data storage devices, network switches, software, computer peripherals including printers and webcam ...
, 1956)
* '' Amalda booleyi'' (Melvill & Sykes, 1896) * '' Amalda borshengi'' Lan & Lee, 2002 * † '' Amalda brevicula'' (Marwick, 1931) * † '' Amalda bruneiana'' Harzhauser, Raven & Landau, 2018 * '' Amalda bulbosa'' Ninomiya, 1991 * '' Amalda bullioides'' (Reeve, 1864) * '' Amalda cacao'' Kantor, Castelin, Fedosov & Bouchet, 2020 * † '' Amalda cincta'' (P. Marshall, 1918) * † '' Amalda cingulata'' (Olson, 1956) * '' Amalda coccinata'' Kilburn, 1980 * '' Amalda coenobium'' Ninomiya, 1991 * '' Amalda colemani'' Ninomiya, 1991 * '' Amalda concinna'' Ninomiya, 1990 * '' Amalda contusa'' (Reeve, 1864) * '' Amalda coriolis'' Kilburn & Bouchet, 1988 * '' Amalda crosnieri'' Kilburn, 1977 * '' Amalda cupedula'' Kilburn, 1993 * '' Amalda danilai'' Kilburn, 1996 * '' Amalda decipiens'' (G. B. Sowerby III, 1897) * '' Amalda degalleaniae'' Cossignani, 2018 * '' Amalda depressa'' (G.B. Sowerby II, 1859) * '' Amalda dimidiata'' (G.B. Sowerby II, 1859) * † '' Amalda dubia'' (Deshayes, 1830) * '' Amalda edgariana'' Schepman, 1911 * '' Amalda edithae'' (Pritchard & Gabriel, 1898) * † '' Amalda elongata'' (Deshayes, 1830) * † '' Amalda exsputa'' (Bartrum & Powell, 1928) * '' Amalda fasciata'' Ninomiya, 1991 * '' Amalda festiva'' Ninomiya, 1991 * '' Amalda fuscolingua'' Kilburn & Bouchet, 1988 * '' Amalda gabelishi'' Ninomiya, 1988 * † '' Amalda garsensis'' Lozouet, 2011 * † '' Amalda gigartoides'' (Olson, 1956) * † '' Amalda glandiformis'' (Lamarck, 1811) * † '' Amalda gulosa'' (C. A. Fleming, 1943) * '' Amalda harasewychi'' Thach, 2016 * '' Amalda hayashii'' Ninomiya, 1988 * '' Amalda herberti'' Cossignani, 2013 * '' Amalda hilgendorfi'' (Martens, 1897) * '' Amalda hinomotoensis'' (Yokoyama, 1922) * † '' Amalda ivrigleyi'' (Chavan & M. Fischer, 1939) * '' Amalda jenneri'' Kilburn, 1977 * '' Amalda josecarlosi'' Pastorino, 2003 * '' Amalda lactea'' T. Kuroda, 1960 * '' Amalda lematrei'' Kilburn, 1977 * † '' Amalda ligata'' (Tate, 1889) * '' Amalda lindae'' Kilburn, 1993 * '' Amalda lineata'' (Kiener, 1844) * '' Amalda lochii'' Ninomiya, 1990 * † '' Amalda macbeathi'' (Vella, 1954) * † '' Amalda malacitana'' Vera-Peláez, 2022 * '' Amalda mamillata'' (Hinds, 1844) * '' Amalda marginata'' (
Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biolo ...
, 1811)
* '' Amalda mirabelflorenti'' Cossignani, 2018 * † '' Amalda miriensis'' Raven & Recourt, 2018 * '' Amalda miriky'' Kantor, Castelin, Fedosov & Bouchet, 2020 * '' Amalda monilifera'' (Reeve, 1864) * '' Amalda montrouzieri'' (Souverby, 1860) * † '' Amalda morgani'' (R. S. Allan, 1926) * '' Amalda mozambicana'' Rosado & Monteiro, 2023 * '' Amalda mucronata'' (G.B. Sowerby I, 1830) * '' Amalda nitidanosum'' Ninomiya, 1991 * '' Amalda northlandica'' (Hart, 1995) * '' Amalda novaezelandiae'' (G.B. Sowerby II, 1859) * '' Amalda obesa'' (G.B. Sowerby II, 1859) * '' Amalda oblonga'' (G.B. Sowerby I, 1830) * † '' Amalda obsoleta'' (Brocchi, 1814) * '' Amalda obtusa'' (Swainson, 1825) * † '' Amalda olivula'' (Lamarck, 1803) * † '' Amalda olsoni'' Beu, 1970 * † '' Amalda opima'' (Marwick, 1924) * '' Amalda optima'' (G.B. Sowerby III, 1897) * † '' Amalda oraria'' (Olson, 1956) * '' Amalda ornata'' Ninomiya, 1988 * '' Amalda otohime'' Majima, Tsuchida & Oshima, 1993 * † '' Amalda ovalis'' Beu, 1970 * '' Amalda pacei'' Petuch, 1987 * † '' Amalda pakaurangiensis'' (Olson, 1956) * '' Amalda parentalis'' Shikama & Oishi, 1977 * † '' Amalda patula'' (Bellardi, 1882) * '' Amalda petterdi'' (Tate, 1893) * '' Amalda pinguis'' Ninomiya, 1991 * † '' Amalda platycephala'' (Powell & Bartrum, 1929) * † '' Amalda pliopenis'' Vera-Peláez, 2022 * '' Amalda ponderi'' Ninomiya, 1991 * † '' Amalda pristina'' (Olson, 1956) * '' Amalda procera'' Ninomiya, 1991 * '' Amalda pullarium'' Ninomiya, 1991 * '' Amalda raoulensis'' Powell, 1967 * '' Amalda reevei'' (E.A. Smith, 1904) * † '' Amalda rimuensis'' (Olson, 1956) * † '' Amalda robusta'' (Marwick, 1924) * '' Amalda roscoae'' Kilburn, 1975 * '' Amalda rottnestensis'' Ninomiya, 1991 * '' Amalda rubiginosa'' (Swainson, 1823) * '' Amalda rubrofasciata'' Ninomiya, 1991 * † '' Amalda schmidti'' (Beets, 1984) * '' Amalda scopuloceti'' Kilburn, 1993 * † '' Amalda semilaevis'' (Tenison Woods, 1879) * '' Amalda similis'' (G.B. Sowerby II, 1859) * '' Amalda sinensis'' (G.B. Sowerby II, 1859) * '' Amalda southlandica'' (Fleming, 1948) * † '' Amalda spinigera'' (P. Marshall, 1918) * † '' Amalda stortha'' (Olson, 1956) * † '' Amalda subampliata'' (Tate, 1889) * † '' Amalda subgradata'' (Tate, 1889) * † '' Amalda subulata'' (Lamarck, 1803) * '' Amalda tankervillii'' (Swainson, 1825) * '' Amalda telaaraneae'' Kilburn, 1993 * '' Amalda tenuis'' Ninomiya, 1991 * † '' Amalda tholiculus'' (Marwick, 1931) * '' Amalda tindalli'' (Melvill, 1898) * † '' Amalda tirangiensis'' (Marwick, 1926) * '' Amalda trachyzonus'' Kilburn, 1975 * '' Amalda trippneri'' Kilburn, 1996 * '' Amalda turgida'' Ninomiya, 1990 * '' Amalda utopica''Ninomiya, 1987 * '' Amalda venezuela'' Weisbord, 1962 * '' Amalda vernedei'' (G.B. Sowerby II, 1859) * '' Amalda virginea'' Ninomiya, 1990 * † '' Amalda waikaiaensis'' (Finlay, 1926) * † ''
Amalda wairarapaensis ''Amalda'' is a genus of medium-sized sea snails, a marine (ocean), marine gastropod mollusc in the family Ancillariidae, the olives and allies.Bouchet, P. (2014). Amalda H. Adams & A. Adams, 1853. Accessed through: World Register of Marine Speci ...
'' (Olson, 1956) * '' Amalda whatmoughi'' Kilburn, 1993


Species brought into synonymy

* ''Amalda albicallosa'' ic synonym of ''Amalda albocallosa'' (Lischke, 1873) * ''Amalda ampla'' (Gmelin, 1791): synonym of '' Ancilla ampla'' (Gmelin, 1791) * † ''Amalda buccinoides'' (Lamarck, 1803): synonym of † '' Spirancilla buccinoides'' (Lamarck, 1803) * ''Amalda callifera'' Thiele, 1925: synonym of ''Amalda reevei'' (E.A. Smith, 1904) * ''Amalda dyspetes'' (Iredale, 1924): synonym of '' Amalda marginata'' (Lamarck, 1811) * ''Amalda elongata'' (Gray, 1874): synonym of '' Ancillista muscae'' (Pilsbry, 1926) * ''Amalda errorum'' Tomlin, 1921: synonym of ''Amalda angustata'' (G.B. Sowerby II, 1859) * ''Amalda fusiformis'' (Petterdi, 1886): synonym of '' Amalda petterdi'' (Tate, 1893) (invalid: not J. Sowerby, 1850) * ''Amalda hayashi'' Ninomiya, 1988: synonym of ''Amalda rubiginosa'' (Swainson, 1823) * ''Amalda lanceolata'' Ninomiya, 1991: synonym of Amalda petterdi ''(Tate, 1893)'' (junior subjective synonym) * † ''Amalda lanceolata'' (Tate, 1889): synonym of † '' Baryspira lanceolata'' (Tate, 1889) (superseded combination) * ''Amalda maritzae'' Bozzetti, 2007: synonym of '' Amalda allaryi'' Bozzetti, 2007 (uncertain synonym) * ''Amalda procerum'' Ninomiya, 1991: synonym of ''Amalda procera'' Ninomiya, 1991 * ''Amalda siberutensis'' Thiele, 1925: synonym of '' Ancilla siberutensis'' Thiele, 1925 * ''Amalda sibuetae'' Kantor & Bouchet, 1999: synonym of '' Turrancilla sibuetae'' (Kantor & Bouchet, 1999) (original combination) * ''Amalda sidneyensis'' Ninomiya, 1991: synonym of '' Amalda edithae'' (Pritchard & Gatliff, 1899) (junior subjective synonym) * ''Amalda tankervillei'' (Swainson, 1825): synonym of ''Amalda tankervillii'' (Swainson, 1825) * ''Amalda virgineus'' Ninomiya, 1990: synonym of ''Amalda virginea'' Ninomiya, 1990 * ''Amalda zeigleri'' Ninomiya, 1987: synonym of ''Amalda pacei'' Petuch, 1987


References


Further reading

* Powell A. W. B., ''New Zealand Mollusca'', William Collins Publishers Ltd, Auckland, New Zealand 1979 * Kilburn, R.N. & Bouchet, P. 1988. ''The genus Amalda in New Caledonia (Mollusca, Gastropoda, Olividae, Ancillinae)''. Bulletin du Muséum National d'Histoire Naturelle. Paris 4 10: 277-300 * Wilson, B. 1994. ''Australian Marine Shells. Prosobranch Gastropods.'' Kallaroo, WA : Odyssey Publishing Vol. 2 370 pp.


External links


Gray, J. E. (1857). Guide to the Systematic Distribution of Mollusca in the British Museum. Part. I. London pp. XII. 230: pp

Adams H. & Adams A. (1853-1858). The genera of Recent Mollusca; arranged according to their organization. London, van Voorst. Vol. 1: xl + 484 pp.; vol. 2: 661 pp.; vol. 3: 138 pls. [Published in parts: Vol. 1: i-xl (1858), 1–256 (1853), 257–484 (1854). Vol. 2: 1-92 (1854), 93–284 (1855), 285–412 (1856), 413–540 (1857), 541–661 (1858). Vol. 3: pl. 1-32 (1853), 33–96 (1855), 97–112 (1856), 113–128 (1857), 129–138 (1858)

Morgan-Richards (2019) Punctuated equilibrium in New Zealand.
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