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Alvar Palmgren (28 April 1880 – 30 November 1960) was a Finnish
botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
and plant
ecologist Ecology () is the natural science of the relationships among living organisms and their environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere levels. Ecology overlaps with the closely re ...
. Palmgren studied
botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
at the
University of Helsinki The University of Helsinki (, ; UH) is a public university in Helsinki, Finland. The university was founded in Turku in 1640 as the Royal Academy of Åbo under the Swedish Empire, and moved to Helsinki in 1828 under the sponsorship of Alexander ...
under professor J.P. Norrlin. He graduated (Fil.kand.) in 1906 and obtained his Ph.D. in 1914. He became docent of
botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
at the University of Helsinki 1916 and professor of
botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
at the same university in 1928 (from 1938, the first in the special
Swedish Swedish or ' may refer to: Anything from or related to Sweden, a country in Northern Europe. Or, specifically: * Swedish language, a North Germanic language spoken primarily in Sweden and Finland ** Swedish alphabet, the official alphabet used by ...
-language chair of botany). He retired in 1950.


Research

Palmgren worked as a
systematic Systematic may refer to: Science * Short for systematic error * Systematic fault * Systematic bias, errors that are introduced by an inaccuracy inherent to the system Economy * Systematic trading, a way of defining trade goals, risk control ...
botanist with
microspecies In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each ot ...
of ''
Taraxacum ''Taraxacum'' () is a genus of flowering plants in the family Asteraceae, which consists of species commonly known as dandelions. The scientific and hobby study of the genus is known as taraxacology. The genus has a near-cosmopolitan distribu ...
'', ''
Hieracium ''Hieracium'' (), known by the common name hawkweed and classically as (from ancient Greek ἱέραξ, 'hawk'), is a genus of flowering plant in the family Asteraceae, and closely related to dandelion (''Taraxacum''), chicory (''Cichorium''), ...
'' and other
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
. As an ecologist, he worked of the nature of
plant communities A plant community is a collection or Association (ecology), association of plant species within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different vegetation types. The comp ...
. He supported the ideas of
Henry Gleason Henry Allan Gleason (1882–1975) was an American ecologist, botanist, and taxonomist. He was known for his endorsement of the individualistic or open community concept of ecological succession, and his opposition to Frederic Clements's concept ...
on the individualistic behaviour of species in community assembly already from the 1920s. Palmgren wrote early accounts on the role of isolation and
stochastic Stochastic (; ) is the property of being well-described by a random probability distribution. ''Stochasticity'' and ''randomness'' are technically distinct concepts: the former refers to a modeling approach, while the latter describes phenomena; i ...
events in the distribution of species, while his contemporary biology was largely
deterministic Determinism is the metaphysical view that all events within the universe (or multiverse) can occur only in one possible way. Deterministic theories throughout the history of philosophy have developed from diverse and sometimes overlapping mo ...
.Palmgren, A. (1921) Die Entfernung als pflanzengeographischer Faktor (Isolation as a phytogeographical factor). Series Acta Societatis pro Fauna et Flora Fennica 49 (1)


Jaccard dispute

In the 1920s, Palmgren entered in a heated dispute with the
Swiss Swiss most commonly refers to: * the adjectival form of Switzerland * Swiss people Swiss may also refer to: Places * Swiss, Missouri * Swiss, North Carolina * Swiss, West Virginia * Swiss, Wisconsin Other uses * Swiss Café, an old café located ...
botanist and
phytogeographer Phytogeography (from Greek φυτόν, ''phytón'' = "plant" and γεωγραφία, ''geographía'' = "geography" meaning also distribution) or botanical geography is the branch of biogeography that is concerned with the geographic distribution o ...
Paul Jaccard Paul Jaccard (18 November 1868 in Sainte-Croix, Switzerland, Sainte-Croix – 9 May 1944 in Zurich) was a professor of botany and plant physiology at the ETH Zurich. He studied at the University of Lausanne and ETH Zurich (PhD 1894). He continued s ...
over the interpretation of Jaccard's species-to-genus ratio. Palmgren had observed a decrease in species richness from west to east in
Åland Åland ( , ; ) is an Federacy, autonomous and Demilitarized zone, demilitarised region of Finland. Receiving its autonomy by a 1920 decision of the League of Nations, it is the smallest region of Finland by both area () and population (30,54 ...
, his main geographical scene of scientific inquiry. He interpreted this as an effect of isolation from the
Swedish Swedish or ' may refer to: Anything from or related to Sweden, a country in Northern Europe. Or, specifically: * Swedish language, a North Germanic language spoken primarily in Sweden and Finland ** Swedish alphabet, the official alphabet used by ...
mainland to the west, and the associated lower species-to-genus ratio as a random sampling effect. In contrast, Jaccard held that the lower species-to-genus ratio towards the east was an effect of decreased diversity in habitat conditions and increased competitive exclusion. The
Swiss Swiss most commonly refers to: * the adjectival form of Switzerland * Swiss people Swiss may also refer to: Places * Swiss, Missouri * Swiss, North Carolina * Swiss, West Virginia * Swiss, Wisconsin Other uses * Swiss Café, an old café located ...
botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
Arthur Maillefer showed statistically that
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
accumulate much faster than
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
and that therefore Jaccard's biological explanation of the pattern was unnecessary since it could be fully explained as a statistical sampling effect. An analytical solution with the same result was provided by the Hungarian
mathematician A mathematician is someone who uses an extensive knowledge of mathematics in their work, typically to solve mathematical problems. Mathematicians are concerned with numbers, data, quantity, mathematical structure, structure, space, Mathematica ...
George Pólya George Pólya (; ; December 13, 1887 – September 7, 1985) was a Hungarian-American mathematician. He was a professor of mathematics from 1914 to 1940 at ETH Zürich and from 1940 to 1953 at Stanford University. He made fundamental contributi ...
. In essence, this disagreement were repeated by
Charles Elton Charles Elton may refer to: * Charles Isaac Elton (1839–1900), English lawyer, politician, writer and antiquarian * Charles Sutherland Elton (1900–1991), English biologist * Charles Elton (police), Chief of Police in Los Angeles, California (19 ...
vs. C. B. Williams and again reiterated by Peter Grant and
Daniel Simberloff Daniel Simberloff is an American biologist and ecologist. He earned his Ph.D. from Harvard University in 1969. He is currently Gore Hunger Professor of Environmental Science at the University of Tennessee, editor-in-chief of the journal ''Bio ...
in the 1970s.


Miscellaneous

Palmgren was on the board of
Societas pro Fauna et Flora Fennica ''Consensu'' or ''obligatio consensu'' or ''obligatio consensu contracta'' or ''obligations ex consensu''George Bowyer, ''Commentaries on the Modern Civil Law'' (London: V & R Stevens and G S Norton, 1848), chapter 26p 201 or ''contractus ex cons ...
from 1916. He was chairman 1920–1957. Palmgren (co-) edited two
exsiccata Exsiccata (Latin, ''gen.'' -ae, ''plur.'' -ae) is a work with "published, uniform, numbered set of preserved specimens distributed with printed labels". Typically, exsiccatae are numbered collections of dried herbarium Biological specimen, spe ...
works, namely ''Carices fulvellae Fries'' and together with L. Fagerström ''Carices extensae exsiccatae a Museo Botanico Universitatis Helsingiensis distributae''. In addition to his botanical work, Palmgren made an impressive political act as a 22-year-old student. He initiated and led a movement among young Finnish men to refuse
conscription Conscription, also known as the draft in the United States and Israel, is the practice in which the compulsory enlistment in a national service, mainly a military service, is enforced by law. Conscription dates back to antiquity and it conti ...
service in the
Russian Imperial The Russian Empire was an empire that spanned most of northern Eurasia from its establishment in November 1721 until the proclamation of the Russian Republic in September 1917. At its height in the late 19th century, it covered about , roughl ...
Army. A rescript of 1900 by
Nicholas II of Russia Nicholas II (Nikolai Alexandrovich Romanov; 186817 July 1918) or Nikolai II was the last reigning Emperor of Russia, Congress Poland, King of Congress Poland, and Grand Duke of Finland from 1 November 1894 until Abdication of Nicholas II, hi ...
, sovereign of the
Grand Duchy of Finland The Grand Duchy of Finland was the predecessor state of modern Finland. It existed from 1809 to 1917 as an Autonomous region, autonomous state within the Russian Empire. Originating in the 16th century as a titular grand duchy held by the Monarc ...
, put Finnish
conscripts Conscription, also known as the draft in the United States and Israel, is the practice in which the compulsory enlistment in a national service, mainly a military service, is enforced by law. Conscription dates back to antiquity and it conti ...
under the
Russian Russian(s) may refer to: *Russians (), an ethnic group of the East Slavic peoples, primarily living in Russia and neighboring countries *A citizen of Russia *Russian language, the most widely spoken of the Slavic languages *''The Russians'', a b ...
military top. The 'strike' movement forced a withdrawal by 1905. Alvar Palmgren was the father of the zoologist
Pontus Palmgren Pontus Palmgren (27 April 1907 – 26 November 1993) was a Finnish ornithologist, arachnologist, and a professor of zoology at the University of Helsinki. He pioneered ethology and quantitative ecology in Finland. Life and work Palgren was born ...
(1907–1993).


References


External links


Swedish biography with portrait
{{DEFAULTSORT:Palmgren, Alvar 20th-century Finnish botanists Finnish ecologists Academic staff of the University of Helsinki 1880 births 1960 deaths Plant ecologists