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''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)
Molecular phylogenetics of ''Alternanthera'' (Gomphrenoideae, Amaranthaceae): resolving a complex taxonomic history caused by different interpretations of morphological characters in a lineage with C4 and C3–C4 intermediate species.
''Botanical Journal of the Linnean Society'' 169(3), 493-517.
''Alternanthera''.
Flora of North America.
and others in Asia, Africa, and Australia. Plants of the genus may be known generally as joyweeds, or Joseph's coat.''Alternanthera''.
Integrated Taxonomic Information System (ITIS).
Several species are notorious
noxious weed A noxious weed, harmful weed or injurious weed is a weed that has been designated by an agricultural or other governing authority as a plant that is injurious to agricultural or horticultural crops, natural habitats or ecosystems, or humans or liv ...
s.Tanveer, A., et al. (2013)
A review on genus ''Alternanthera'' weeds implications.
''Pak. J. Weed Sci. Res'' 19(1), 53-58.


Description

These are annual or perennial herbs or
subshrub A subshrub (Latin ''suffrutex'') or dwarf shrub is a short shrub, and is a woody plant. Prostrate shrub is a related term. "Subshrub" is often used interchangeably with "bush".Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their Der ...
s. While some of the better-known species are
aquatic plant Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...
s, most are terrestrial. They take many forms, from prostrate to erect to floating. The leaves are oppositely arranged. The inflorescence is a spike or a rounded head occurring in the leaf axils or the ends of branches. The flowers have 5 tepals. There are 3 to 5
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s which are fused into a rim at the bases, and 5 pseudostaminodes, appendages between the stamens that are not true
staminode In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co. ...
s. The fruit is a utricle containing one seed. The genus ''Alternanthera'' contains both terrestrial and aquatic species. The
photosynthetic Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in c ...
pathway varies in this genus: Some species undergo C3 carbon fixation, one clade of 17 species has acquired the C4 pathway, and yet other species have an intermediate C3-C4 pathway. Chinthapalli, B., et al. (2001)
Phospho''enol''pyruvate carboxylase purified from leaves of C3, C4, and C3-C4 intermediate species of ''Alternanthera'': Properties at limiting and saturating bicarbonate.
''Photosynthetica'' 38(3), 415-19.


Species

It is not yet clear how many species belong in the genus. Estimates range between 80 and 200.''Alternanthera''.
The Plant List.
Species include:GRIN Species Records of ''Alternanthera''.
Germplasm Resources Information Network (GRIN).
*'' Alternanthera acaulis'' Andersson *''
Alternanthera adscendens ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomphren ...
'' Suess. *''
Alternanthera albotomentosa ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomphren ...
'' Suess. *''
Alternanthera albida ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomphren ...
'' (Moq.) Griseb *''
Alternanthera albosquarrosa ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomphren ...
'' Suess. *''
Alternanthera albotomentosa ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomphren ...
'' Suess. *''
Alternanthera altacruzensis ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomphren ...
'' Suess. *'' Alternanthera angustifolia'' R.Br. – narrow-leaf joyweed *''
Alternanthera aquatica ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomphren ...
'' (Parodi) Chodat *''
Alternanthera arequipensis ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomphren ...
'' Suess. *''
Alternanthera areschougii ''Alternanthera areschougii'' is a species of plant in the family Amaranthaceae. It is endemic to Ecuador. Its natural habitats are subtropical or tropical dry forests and subtropical or tropical dry shrubland Shrubland, scrubland, scr ...
'' *''
Alternanthera asterotricha ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomph ...
'' Uline *''
Alternanthera axillaris ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomph ...
'' (Willd.) DC. *''
Alternanthera bettzickiana ''Alternanthera bettzickiana'', commonly known as calico-plant, is a species of flowering plant in the family Amaranthaceae. It is commonly used as an ornamental edging plant. Native to South America, its cultivar 'Red' is similar in appearance ...
'' – calico-plant *''
Alternanthera brasiliana ''Alternanthera brasiliana'', also known as large purple alternanthera, metal weed, bloodleaf, parrot leaf, ruby leaf, Brazilian joyweed, purple alternanthera, purple joyweed, is a flowering plant of the amaranth family that is native to the fore ...
'' (L.) Kuntze – Brazilian joyweed, ruby leaf *'' Alternanthera caracasana'' Kunth – mat chaff-flower, washerwoman *''
Alternanthera corymbiformis ''Alternanthera corymbiformis'' is a species of plant in the family Amaranthaceae. It is endemic to Ecuador. Its natural habitats are subtropical or tropical moist montane forests and subtropical or tropical dry shrubland Shrubland, sc ...
'' *''
Alternanthera crucis ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomphren ...
'' Bold. – West Indian joyweed *''
Alternanthera dentata ''Alternanthera dentata'', known as little ruby and ruby leaf alternanthera, is a fast-growing ornamental groundcover plant in the amaranth family which was first described by Conrad Moench, and got its current name from Stuchlík and Robert Elia ...
'' – little ruby *''
Alternanthera denticulata ''Alternanthera denticulata'' (common name lesser joyweed) is a small prostrate white-flowering herb in the Amaranthaceae family. It is native to all states and territories of Australia, New Guinea, and the North Island The North Island, ...
'' – lesser joyweed *''
Alternanthera echinocephala ''Alternanthera echinocephala'', known as spiny-headed chaff flower, is a shrubby plant in the family Amaranthaceae native to the Galápagos Islands, mainland Ecuador, and Peru. Its relatively large spiny "heads" of flowers distinguish it fr ...
'' (Hook.f.) Christoph. – spiny-headed chaff flower,, p. 144 sea-urchin joyweed *''
Alternanthera ficoidea ''Alternanthera ficoidea'', also known as Joseph's coat, Parrot leaf, is a species of flowering plant in the family Amaranthaceae. It is native to the Caribbean and South America and occurs elsewhere in the tropics as an introduced species. I ...
'' – parrot leaf, sanguinarea *''
Alternanthera filifolia ''Alternanthera filifolia'' is a species of flowering plant in the family Amaranthaceae. It is native to the Galápagos Islands. This plant is limited to the Galápagos, but it is common there in lowlands and coastal shrublands. It is variable i ...
'' (Hook.f.) J.T. Howell *''
Alternanthera flavescens ''Alternanthera'' is a genus of flowering plants in the family Amaranthaceae. It is a widespread genus with most species occurring in the tropical Americas,Sánchez-Del Pino, I., et al. (2012)Molecular phylogenetics of ''Alternanthera'' (Gomph ...
'' – yellow joyweed *''
Alternanthera flavicoma ''Alternanthera flavicoma'' is a species of plant in the family Amaranthaceae. It is endemic to Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuad ...
'' *''
Alternanthera galapagensis ''Alternanthera galapagensis'' is a species of plant in the family Amaranthaceae. It is endemic to the Galapagos. References Flora of Ecuador galapagensis Vulnerable plants Taxonomy articles created by Polbot Plants described in 191 ...
'' *''