Allotheria (meaning "other beasts", from the
Greek , '–other and , '–wild animal) is an extinct branch of successful
Mesozoic
The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceo ...
mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s. The most important characteristic was the presence of lower
molariform teeth
A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
equipped with two longitudinal rows of cusps. Allotheria includes
Multituberculata,
Gondwanatheria
Gondwanatheria is an extinct group of mammaliaforms that lived in parts of Gondwana, including Madagascar, India, South America, Africa and Antarctica during the Upper Cretaceous through the Paleogene (and possibly much earlier, if '' Allostaff ...
(which may be part of
Multituberculata, as the sister group to
Cimolodonta),
and probably
Haramiyida
Haramiyida ("thief" from Arabic الحرامية (al ḥarāmiyah), "thief, bandit") is a possibly polyphyletic order of mammaliaform cynodonts or mammals of controversial taxonomic affinites. Their teeth, which are by far the most common remains ...
, although some studies show them to be more basal
mammaliaforms rather than true mammals, therefore differing from true allotheres significantly.
Allotheres also had a narrow pelvis, indicating that they gave birth to tiny helpless young like marsupials or laid eggs and gave milk to feed their young like
monotreme
Monotremes () are prototherian mammals of the order Monotremata. They are one of the three groups of living mammals, along with placentals (Eutheria), and marsupials (Metatheria). Monotremes are typified by structural differences in their brain ...
s do. This is a feature of all non-
placental mammals.
Interpretations
When he first identified Allotheria in 1880,
Othniel Marsh regarded this group as an order within
Marsupialia. However, in 1997 McKenna and Bell classified Allotheria as an
infraclass
In biological classification, class ( la, classis) is a taxonomic rank, as well as a taxonomic unit, a taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are life, domain, kingdom ...
.
References
Further reading
Zofia Kielan-Jaworowska, Richard L. Cifelli, and Zhe-Xi Luo, ''Mammals from the Age of Dinosaurs: Origins, Evolution, and Structure'' (New York: Columbia University Press, 2004), 249.
{{Taxonbar, from=Q1419784
Prehistoric mammals
Mammal taxonomy
Norian first appearances
Late Triassic first appearances
Miocene extinctions
Fossil taxa described in 1880