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''Albicetus'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
stem Stem or STEM most commonly refers to: * Plant stem, a structural axis of a vascular plant * Stem group * Science, technology, engineering, and mathematics Stem or STEM can also refer to: Language and writing * Word stem, part of a word respon ...
-
sperm whales The sperm whale or cachalot (''Physeter macrocephalus'') is the largest of the toothed whales and the largest toothed predator. It is the only living member of the genus '' Physeter'' and one of three extant species in the sperm whale super ...
that lived during the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
Epoch, around 15 million years ago, and was discovered in
Santa Barbara, California Santa Barbara (, meaning ) is a coastal city in Santa Barbara County, California, of which it is also the county seat. Situated on a south-facing section of coastline, the longest such section on the West Coast of the United States excepting A ...
in 1909. It was categorized for decades as belonging to a group of extinct
walrus The walrus (''Odobenus rosmarus'') is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. It is the only extant species in the family Odobeni ...
es erroneously thought to be sperm whales. It was named ''Albicetus'', meaning "white whale", is a reference to the leviathan in
Herman Melville Herman Melville (Name change, born Melvill; August 1, 1819 – September 28, 1891) was an American novelist, short story writer, and poet of the American Renaissance (literature), American Renaissance period. Among his best-known works ar ...
's classic 1851 novel ''
Moby-Dick ''Moby-Dick; or, The Whale'' is an 1851 Epic (genre), epic novel by American writer Herman Melville. The book is centered on the sailor Ishmael (Moby-Dick), Ishmael's narrative of the maniacal quest of Captain Ahab, Ahab, captain of the whaler ...
''.


Taxonomy


Etymology

The genus ''Albicetus'' derives from the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
''albus'' "white" and ''cetus'' "whale", and so literally means "white whale". It was named in reference to the
antagonist An antagonist is a character in a story who is presented as the main enemy or rival of the protagonist and is often depicted as a villain.Moby Dick ''Moby-Dick; or, The Whale'' is an 1851 Epic (genre), epic novel by American writer Herman Melville. The book is centered on the sailor Ishmael (Moby-Dick), Ishmael's narrative of the maniacal quest of Captain Ahab, Ahab, captain of the whaler ...
from
Herman Melville Herman Melville (Name change, born Melvill; August 1, 1819 – September 28, 1891) was an American novelist, short story writer, and poet of the American Renaissance (literature), American Renaissance period. Among his best-known works ar ...
's novel ''
Moby-Dick ''Moby-Dick; or, The Whale'' is an 1851 Epic (genre), epic novel by American writer Herman Melville. The book is centered on the sailor Ishmael (Moby-Dick), Ishmael's narrative of the maniacal quest of Captain Ahab, Ahab, captain of the whaler ...
''. This reference was done both in honor of Melville and as a reference to the most notable traits of the Moby Dick–including an "unwonted magnitude", light color, and a crooked and deformed lower jaw–which coincidentally also describe the traits found in the ''Albicetus oxymycterus''
holotype specimen A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was Species description, formally described. It is either the single such physical example (or illus ...
(the fossils themselves possess a light hue). The species name ''oxymycterus'' is derived from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
ὀξύς ''oxy'' "sharp" and μυκτήρ ''mycter'' "nose".


Taxonomic history

The holotype was recovered from an unspecified location in the sea cliffs near the original Santa Barbara Lighthouse in 1909, which is believed to most likely be part of the
Monterey Formation The Monterey Formation is an extensive Miocene oil-rich geology, geological sedimentary formation in California, with outcrops of the formation in parts of the California Coast Ranges, Peninsular Ranges, and on some of California's off-shor ...
. The whale was originally placed in the genus ''
Ontocetus ''Ontocetus'' is an extinct genus of walrus, an aquatic carnivoran of the family Odobenidae, endemic to coastal regions of the southern North Sea and the southeastern coastal regions of the U.S. during the Miocene-Pleistocene. It lived from 1 ...
'' in 1925 by American naturalist
Remington Kellogg Arthur Remington Kellogg (5 October 1892 – 8 May 1969) was an American naturalist and a director of the United States National Museum. His work focused on marine mammals. Early life and career Kellogg was born in Davenport, Iowa, and quickly ...
as ''O. oxymycterus''. This genus was originally thought to represent a sperm whale, however, in 2008, the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
, ''Ontocetus emmonsi'', was discovered to actually be a
walrus The walrus (''Odobenus rosmarus'') is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. It is the only extant species in the family Odobeni ...
. The whale was then moved to the
wastebasket taxon Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon) is a term used by some taxonomists to refer to a taxon that has the purpose of classifying organisms that do not fit anywhere else. They are typically defined by e ...
''
Scaldicetus ''Scaldicetus'' is an extinct genus of highly predatory macroraptorial sperm whale. Although widely used for a number of extinct physeterids with primitive dental morphology consisting of enameled teeth, ''Scaldicetus'' as generally recognized ...
'', which consists of various other (more-or-less unrelated) primitive sperm whales with enamel coated teeth. In 2015, the whale was moved to the newly erected genus ''Albicetus''.


Phylogeny

''Albicetus'', unlike the modern
sperm whale The sperm whale or cachalot (''Physeter macrocephalus'') is the largest of the toothed whales and the largest toothed predator. It is the only living member of the Genus (biology), genus ''Physeter'' and one of three extant species in the s ...
(''Physeter macrocephalus''), possessed functional and enamel-coated teeth in both jaws. This suggests that it is related to the group of
macroraptorial sperm whale Macroraptorial sperm whales were highly predatory whales of the sperm whale superfamily (Physeteroidea) of the Miocene epoch (geology), epoch that hunted large marine mammals, including other whales, using their large teeth. They consist of six ge ...
s which includes '' Acrophyseter'', ''
Brygmophyseter ''Brygmophyseter'', known as the biting sperm whale, is an extinct genus of toothed whale in the Physeteroidea, sperm whale family with one species, ''B. shigensis''. When it was first described in 1994, the species was placed in the genus ''Sc ...
'', ''
Livyatan ''Livyatan'' is an extinct genus of macroraptorial Physeteroidea, sperm whale containing one known species: ''L. melvillei''. The genus name was inspired by the Bible, biblical sea monster Leviathan, and the species name by Herman Melville, th ...
'', and ''
Zygophyseter ''Zygophyseter varolai'' is an extinct sperm whale The sperm whale or cachalot (''Physeter macrocephalus'') is the largest of the toothed whales and the largest toothed predator. It is the only living member of the Genus (biology), genu ...
''. However, ''Albicetus'' is most similar in general shape and characteristics with '' Aulophyseter morricei'', except for the dentition in which the latter possesses only small, vestigial upper teeth lacking enamel. The closest known relative to ''Albicetus'' is currently ''Livyatan''. ''Albicetus'' contains only one species ''A. oxymycterus''.


Description

The
type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
,
USNM The National Museum of Natural History (NMNH) is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. With 4.4 ...
10923, consists of a partial skull (mainly remains of the beak) and isolated tooth fragments. Comparing the
occipital condyle The occipital condyles are undersurface protuberances of the occipital bone in vertebrates, which function in articulation with the superior facets of the Atlas (anatomy), atlas vertebra. The condyles are oval or reniform (kidney-shaped) in shape ...
length with antorbital notch (slits in the skull right before the snout) width of other primitive sperm whale, the total body length is estimated to be between . The preserved length of the beak is . The whale had a maximum of 18 teeth in either jaw, deeply rooted in exceedingly large
tooth socket Dental alveoli (singular ''alveolus'') are sockets in the jaws in which the roots of teeth are held in the alveolar process with the periodontal ligament. The lay term for dental alveoli is tooth sockets. A joint that connects the roots of the t ...
s.


Paleoecology

For macroraptorial sperm whales, the presence of large body size along with large tooth size suggests that body size was an adaptation to hunting bigger prey, notably other
marine mammal Marine mammals are mammals that rely on marine ecosystems for their existence. They include animals such as cetaceans, pinnipeds, sirenians, sea otters and polar bears. They are an informal group, unified only by their reliance on marine enviro ...
s, as opposed to the modern sperm whale where size is possibly an adaptation to deep diving. The
Langhian The Langhian is, in the ICS geologic timescale, an age or stage in the middle Miocene Epoch/Series. It spans the time between 15.97 ± 0.05 Ma and 13.65 ± 0.05 Ma (million years ago) during the Middle Miocene.GeoWhen (2007) The Langhian was ...
is particularly rich in sperm whale diversity, with ''Albicetus'', ''Brygmophyseter'', and ''
Aulophyseter ''Aulophyseter'' is an extinct genus of sperm whales from the Miocene formations of the west and east coasts of North America. ''Aulophyseter'' reached a length of approximately with an estimated body weight of .R. Nieuwenhuys (1998) ''The cent ...
'' contemporaneously inhabiting the North Pacific. It is possible these sperm whales exhibited
niche partitioning In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for e ...
to avoid directly competing for food.


References


External links

* {{Taxonbar, from=Q22285172 Sperm whales Prehistoric toothed whales Miocene cetaceans Miocene mammals of North America Monotypic prehistoric cetacean genera Fossil taxa described in 2015 Paleontology in California