Albertonykus Borealis
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''Albertonykus'' (meaning "
Alberta Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
claw") is an
alvarezsaurid Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from the
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-age (
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cret ...
) rocks of the
Horseshoe Canyon Formation The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller. The Horseshoe Canyon Formation is part of th ...
of Alberta,
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
. It is known from forelimb and hindlimb remains from multiple individuals. All but two of the specimens come from a
bonebed A bone bed is any geological stratum or deposit that contains bones of whatever kind. Inevitably, such deposits are sedimentary in nature. Not a formal term, it tends to be used more to describe especially dense collections such as Lagerstätte. ...
dominated by ''
Albertosaurus ''Albertosaurus'' (; meaning "Alberta lizard") is a genus of large tyrannosaurid theropod dinosaur that lived in northwestern North America during the early to middle Maastrichtian age of the Late Cretaceous period, about 71 million yea ...
'', located at the top of Unit 4 of the Horseshoe Canyon Formation,Larson, D. W., Brinkman, D. B., & Bell, P. R. (2010). Faunal assemblages from the upper Horseshoe Canyon Formation, an early Maastrichtian cool-climate assemblage from Alberta, with special reference to the Albertosaurus sarcophagus bonebed This article is one of a series of papers published in this Special Issue on the theme Albertosaurus. Canadian Journal of Earth Sciences, 47(9), 1159-1181. dating to ~68.5 million years ago. ''Albertonykus'' is interpreted as having fed on wood-nesting
termite Termites are a group of detritivore, detritophagous Eusociality, eusocial cockroaches which consume a variety of Detritus, decaying plant material, generally in the form of wood, Plant litter, leaf litter, and Humus, soil humus. They are dist ...
s because the forelimbs appear specialized for digging, but are too short for burrowing. ''Albertonykus'' is the earliest-known
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
n alvarezsaurid. Isolated bones of alvarezsaurids are known from late Maastrichtian rocks in
Montana Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...
and
Wyoming Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
(
USA The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 states and a federal capital district, Washington, D.C. The 48 contiguous ...
). The bones now belonged to a new genus, ''
Trierarchuncus ''Trierarchuncus'' (meaning "captain hook," after its single-clawed hands) is a monotypic genus of alvarezsauridae, alvarezsaurid theropod which includes a single species, ''Trierarchuncus prairiensis'', mainly known from fossils found in deposits ...
''. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''A. borealis'', described by Nick Longrich and Philip Currie in 2009. The specific name (''borealis'') means "north".


Discovery

Until the 1990s it was unknown that dinosaurs like ''Albertonykus'' existed. A chain of new discoveries from Asia and South America exposed the existence of these previously unknown theropod dinosaurs.Brian Switek, ''Smithsonian Magazine'', http://dinosaur.smithsonianmag.com/2008/10/06/a-dinosaur-that-ate-termites-for-breakfast/ North American finds remained rare. The bones of ''Albertonykus'' were discovered through a coincidental association with a much larger theropod dinosaur. In 1910,
paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
Barnum Brown Barnum Brown (February 12, 1873 – February 5, 1963), commonly referred to as Mr. Bones, was an American paleontologist. He discovered the first documented remains of ''Tyrannosaurus'' during a career that made him one of the most famous fossil ...
and his crew found a site filled with the bones of the gigantic
tyrannosaurid Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to fifteen genera, including the eponymous ''Tyrannosaurus''. The exact number of genera ...
''
Albertosaurus sarcophagus ''Albertosaurus'' (; meaning "Alberta lizard") is a genus of large tyrannosaurid theropod dinosaur that lived in northwestern North America during the early to middle Maastrichtian age of the Late Cretaceous period, about 71 million yea ...
''. They unearthed the remains of at least nine different specimens of ''Albertosaurus'' in the single site, but stopped digging after just two weeks.Nick Longrich, ''University of Calgary'', Renewed quarrying in the ''Albertosaurus'' bonebed in Dry Island Provincial Park again produced the remains of over twenty specimens of ''Albertosaurus'', and the quarry has yet to be fully excavated. It is not known why so many tyrannosaurs were found in one place: paleontologist Nick Longrich speculated that the site was part of a predator trap, where trapped prey animals drew predators to their deaths. At the same site, over a dozen arm and leg bones belonging to an unknown species of small theropod were in 2002 excavated by a team of scientist led by
Philip John Currie Philip John Currie (born March 13, 1949) is a Canadian palaeontologist and museum curator who helped found the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta and is now a professor at the University of Alberta in Edmonton. In the ...
of the
University of Alberta The University of Alberta (also known as U of A or UAlberta, ) is a public research university located in Edmonton, Alberta, Canada. It was founded in 1908 by Alexander Cameron Rutherford, the first premier of Alberta, and Henry Marshall Tory, t ...
. The bones were then stored at the
Royal Tyrrell Museum The Royal Tyrrell Museum of Palaeontology (RTMP; often referred to as the Royal Tyrrell Museum) is a palaeontology museum and research facility in Drumheller, Alberta, Canada. The museum was named in honour of Joseph Tyrrell, Joseph Burr Tyrrell, ...
in Alberta.Ken Than, ''National Geographic News''
/ref>Jeanna Bryner, ''MSNBC.com''
/ref> In 2005 Nick Longrich, then a student at the
University of Calgary {{Infobox university , name = University of Calgary , image = University of Calgary coat of arms without motto scroll.svg , image_size = 150px , caption = Coat of arms , former ...
, happened upon the small fossils while comparing ''Albertosaurus'' claws to other dinosaur species. Longrich recognized the fossils as alvarezsaurid, concluding that they represented a new species. Other fragments of ''Albertonykus'' were then recognized in the collections, which had not been identified as a separate species.


Description

''Albertonykus'' is the smallest known
alvarezsauridae Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly ...
ever discovered in North America, measuring long and weighing . Alvarezsaurs typically had slender hind legs, long rigid tails, and unusually short arms that were powerfully built to support a large claw. Although no skull bones of ''Albertonykus'' have been found, related animals from
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
show that they likely had long, slender snouts filled with tiny teeth similar to those of armadillos and anteaters. It is likely that ''Albertonykus'' ate insects, using its large thumb claw to tear open rotten logs to find its prey. As in other alvarezsaurs, the forelimbs of ''Albertonykus'' were specialized for digging, but were too short to permit burrowing. Unfortunately at this time the skeleton of ''Albertonykus'' is not complete, but its Mongolian relatives give us a distinct idea of what the rest of the skeleton would have looked like. Possible prey items were evaluated and compared with the fossil record of social insects.
Ant Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...
s were not an important part of the ecosystem during the Cretaceous, and mound-building termites do not appear until the Eocene. This leaves the possibility that ''Albertonykus'' preyed on wood-nesting termites. This hypothesis was tested by examining
petrified wood Petrified wood (from Ancient Greek meaning 'rock' or 'stone'; literally 'wood turned into stone'), is the name given to a special type of ''fossilized wood'', the fossilized remains of terrestrial plant, terrestrial vegetation. ''Petrifaction ...
from the
Horseshoe Canyon Formation The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller. The Horseshoe Canyon Formation is part of th ...
, where ''Albertonykus'' was found. The wood found there frequently contains borings, which resemble those of termites.


Classification

''Albertonykus'' is a member of an unusual
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
group known as the
Alvarezsauridae Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly ...
and is one of only a few alvarezsaurid fossils that has been found outside of South America and Asia. After a phylogenetic analysis, it was shown that ''Albertonykus'' is a member of the Asian
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
Mononykini Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly ...
, which supported the hypothesis that the alvarezsaurs originated in South America, and then dispersed to Asia through North America. The unearthing of ''Albertonykus'' provided important information into the biology of the Alvarezsauridae.


References


External links


Small 'Dr. Seuss'-like dinosaur dug up in Alberta
news report at CBC.ca {{Taxonbar, from=Q131750 Alvarezsauridae Dinosaur genera Maastrichtian dinosaurs Horseshoe Canyon Formation Dinosaurs of Canada Fossil taxa described in 2009 Taxa named by Philip J. Currie