Albertaceratops Scale
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''Albertaceratops'' (meaning "
Alberta Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
horned face") is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
centrosaurine Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid, a group of large quadrupedal ornithischian dinosaur. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Alber ...
horned A horn is a permanent pointed projection on the head of various animals that consists of a covering of keratin and other proteins surrounding a core of live bone. Horns are distinct from antlers, which are not permanent. In mammals, true horns ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from the middle
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
-age
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cret ...
Oldman Formation The Oldman Formation is a stratigraphic unit of Late Cretaceous (Campanian stage) age that underlies much of southern Alberta, Canada. It consists primarily of sandstones that were deposited in fluvial channel and floodplain environments. It was ...
of
Alberta Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
,
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
.


Description

''Albertaceratops'' is unusual in combining long brow
horns Horns or The Horns may refer to: * Plural of Horn (anatomy) * Plural of Horn (instrument), a group of musical instruments all with a horn-shaped bells * The Horns (Colorado), a summit on Cheyenne Mountain * Horns (novel), ''Horns'' (novel), a dar ...
with an otherwise centrosaurine skull, as centrosaurines normally possess short brow horns. Over its nose was a bony ridge, and on its frill were two large outwardly-projecting hooks. Its size has been estimated at and .


Discovery and naming

''Albertaceratops'' is known from a single complete skull ( TMP.2001.26.1) found in August 2001 and skull and postcranial fragments. A
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analysis carried out by its describer, Michael J. Ryan, found it to be the most basal centrosaurine. Additional specimens were reported from a bonebed in the
Judith River Formation The Judith River Formation is a fossil-bearing geologic formation in Montana, and is part of the Montana Group. It dates to the Late Cretaceous, between 79 and 75.3 million years ago, corresponding to the "Judithian" land vertebrate age. It was ...
of
Montana Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...
, which is equivalent to the Oldman Formation and differentiated only by the Canada–US border. However, further study showed these remains to come from a different centrosaurine, ''
Medusaceratops ''Medusaceratops'' is an extinct genus of Centrosaurinae, centrosaurine ceratopsian dinosaur known from the Late Cretaceous Judith River Formation (middle Campanian stage) of Montana, northern United States. It contains a single species, ''Medusa ...
''. Both ceratopsids lived during the same time period, about 77.5 million years ago.Ryan, Michael J.; Russell, Anthony P., and Hartman, Scott. (2010). "A New Chasmosaurine Ceratopsid from the Judith River Formation, Montana", In: Michael J. Ryan, Brenda J. Chinnery-Allgeier, and David A. Eberth (eds), ''New Perspectives on Horned Dinosaurs: The Royal Tyrrell Museum Ceratopsian Symposium'', Indiana University Press, 656 pp. . The specific name, ''A. nesmoi'', is derived from the name of Cecil Nesmo, a rancher living in Manyberries,
Alberta Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
, a town of less than 100 people located 71 km south of Medicine Hat. The rancher was thus honored in recognition of his efforts to aid
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
hunters.


Classification

The
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
presented below follows a phylogenetic analysis by Chiba ''et al.'' (2017), which included a systematic re-evaluation of ''
Medusaceratops ''Medusaceratops'' is an extinct genus of Centrosaurinae, centrosaurine ceratopsian dinosaur known from the Late Cretaceous Judith River Formation (middle Campanian stage) of Montana, northern United States. It contains a single species, ''Medusa ...
lokii'':


See also

*
Timeline of ceratopsian research This timeline of ceratopsian research is a chronological listing of events in the History of paleontology, history of paleontology focused on the ceratopsians, a group of herbivorous marginocephalian dinosaurs that evolved parrot-like beaks, b ...


References


External links


Drawing of the skull
by Julius T. Csotonyi