The brook floater or swollen wedgemussel, ''Alasmidonta varicosa'', is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of freshwater mussel, an aquatic

bivalve Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class (biology), class of aquatic animal, aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed b ...

mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...

in the family

Unionidae The Unionidae are a Family (biology), family of freshwater mussels, the largest in the order Unionida, the bivalve molluscs sometimes known as river mussels, or simply as unionids. The range of distribution for this family is world-wide. It is a ...

, the river mussels. It measures 25.1 mm to 80.2 mm in length although other research also suggests it rarely exceeds three inches (75 mm).

Distribution

This species is found in

Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...

(New Brunswick and Nova Scotia) and northeastern

United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...

(

Connecticut Connecticut ( ) is a U.S. state, state in the New England region of the Northeastern United States. It borders Rhode Island to the east, Massachusetts to the north, New York (state), New York to the west, and Long Island Sound to the south. ...

Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...

Maine Maine ( ) is a U.S. state, state in the New England region of the United States, and the northeasternmost state in the Contiguous United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Provinces and ...

Maryland Maryland ( ) is a U.S. state, state in the Mid-Atlantic (United States), Mid-Atlantic region of the United States. It borders the states of Virginia to its south, West Virginia to its west, Pennsylvania to its north, and Delaware to its east ...

Massachusetts Massachusetts ( ; ), officially the Commonwealth of Massachusetts, is a U.S. state, state in the New England region of the Northeastern United States. It borders the Atlantic Ocean and the Gulf of Maine to its east, Connecticut and Rhode ...

New Hampshire New Hampshire ( ) is a U.S. state, state in the New England region of the Northeastern United States. It borders Massachusetts to the south, Vermont to the west, Maine and the Gulf of Maine to the east, and the Canadian province of Quebec t ...

New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...

, New York,

North Carolina North Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, South Carolina to the south, Georgia (U.S. stat ...

Pennsylvania Pennsylvania, officially the Commonwealth of Pennsylvania, is a U.S. state, state spanning the Mid-Atlantic (United States), Mid-Atlantic, Northeastern United States, Northeastern, Appalachian, and Great Lakes region, Great Lakes regions o ...

South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...

Vermont Vermont () is a U.S. state, state in the New England region of the Northeastern United States. It borders Massachusetts to the south, New Hampshire to the east, New York (state), New York to the west, and the Provinces and territories of Ca ...

Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...

and

West Virginia West Virginia is a mountainous U.S. state, state in the Southern United States, Southern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States.The United States Census Bureau, Census Bureau and the Association of American ...

); It was formerly found in

Rhode Island Rhode Island ( ) is a state in the New England region of the Northeastern United States. It borders Connecticut to its west; Massachusetts to its north and east; and the Atlantic Ocean to its south via Rhode Island Sound and Block Is ...

and four watersheds in Massachusetts but are now extinct in Rhode Island and almost extinct in Massachusetts. 1897 Research by Arnold Edward Ortmann showed it to be common in the

Delaware Delaware ( ) is a U.S. state, state in the Mid-Atlantic (United States), Mid-Atlantic and South Atlantic states, South Atlantic regions of the United States. It borders Maryland to its south and west, Pennsylvania to its north, New Jersey ...

and Susquehanna Rivers.

Habitat and behavior

This mussel lives in high relief streams, under boulders and in sand. Research has shown that it is highly sensitive to increased thermal temperature. It associates with longnose and eastern blacknose dace,

golden shiner The golden shiner (''Notemigonus crysoleucas'') is a species of freshwater ray-finned fish belonging to the family Leuciscidae. This fish occurs in Eastern North America. It is the sole member of its genus. Commonly used as a bait fish, it is pr ...

, pumpkinseed,

slimy sculpin The slimy sculpin (''Cottus cognatus'') is a freshwater species of fish belonging to the family Cottidae, which is the largest sculpin family. They usually inhabit cold rocky streams or lakes across North America, ranging from the Great Lakes, s ...

and yellow perch.

Survival threats and conservation

The brook floater is sensitive to habitat loss for development, dams and road crossings, pollution, summer droughts, trampling, sedimentation, flow alteration, and low oxygen conditions. Hybridization with elktoe (''Alasmidonta marginata''), a longtime ally, has also been shown to be a threat. Research has also shown the population is highly fragmented, low in density, prone to mortality due to old age and there are also low chances of longevity and viable reproduction. Trematoda ''rhopalocercous cercaria'' is a parasite of the brook floater. Current research shows populations that were large and widespread have declined by 50% to 95% to almost extinct. While the IUCN lists it as Vulnerable, the states of New Jersey, Virginia, Massachusetts and New Hampshire all list it as Endangered, Threatened in Vermont, Maine and New York, Rare/Endangered in Connecticut, Extinct in Rhode Island and "Species of Special Concern" by the federal government.

References

Alasmidonta Bivalves described in 1819 Taxonomy articles created by Polbot {{Unionidae-stub