The heart and dart (''Agrotis exclamationis'') is a

moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...

of the family

Noctuidae The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family (biology), family of moths. Taxonomically, they are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly ...

. The species was first described by

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

in his 1758 10th edition of ''Systema Naturae''. A familiar moth to many, it is considered one of the most common of the European region. It occurs throughout the

Palearctic realm The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. The ...

from Ireland to Japan. This is a quite variable species with forewings ranging from pale to dark brown but always recognizable by the distinctively shaped dark

stigmata Stigmata (, plural of , 'mark, spot, brand'), in Roman Catholicism, Catholicism, are bodily wounds, scars and pain which appear in locations corresponding to the Crucifixion of Jesus, crucifixion Five Holy Wounds, wounds of Jesus in Christian ...

which give it its common name. The

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...

is 35–44 mm. The hindwings are whitish (compared with other common ''Agrotis'' species, the hindwings of this species are usually paler than in

heart and club The heart and club (''Agrotis clavis'') is a moth of the family Noctuidae. It is distributed throughout the Palearctic realm. The common name of this species refers to the supposed shapes of the bold dark Glossary of entomology terms, stigmata ...

but darker than in

turnip moth ''Agrotis segetum'', sometimes known as the turnip moth, is a moth of the family Noctuidae. The species was first described by Michael Denis and Ignaz Schiffermüller in 1775. It is a common European species and it is found in Africa and across ...

). This species usually has a dark area at the front of the

thorax The thorax (: thoraces or thoraxes) or chest is a part of the anatomy of mammals and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main di ...

, visible as a horizontal bar when viewing the moth head on. The differences are not consistent however; they are highly variable in both colour and markings, and identification of atypical or worn examples may prove impossible without examination of genitalia. See Townsend et al. This moth flies at night from May to July and is attracted to light, sometimes in large numbers. It also frequently visits

nectar Nectar is a viscous, sugar-rich liquid produced by Plant, plants in glands called nectaries, either within the flowers with which it attracts pollination, pollinating animals, or by extrafloral nectaries, which provide a nutrient source to an ...

-rich flowers such as ''

Buddleia ''Buddleja'' (; ''Buddleia''; also historically given as ''Buddlea'') is a genus comprising over 140 species of flowering plants endemicity, endemic to Asia, Africa, and the Americas. The generic name bestowed by Carl Linnaeus, Linnaeus posthu ...

'',

ragwort ''Jacobaea vulgaris'', syn. ''Senecio jacobaea'', is a very common wild flower in the family Asteraceae that is native to northern Eurasia, usually in dry, open places, and has also been widely distributed as a weed elsewhere. Common names inc ...

and

red valerian ''Valeriana rubra'' (synonym ''Centranthus ruber''), the red valerian, spur valerian, kiss-me-quick, fox's brush, devil's beard or Jupiter's beard, is a popular garden plant grown for its ornamental flowers. Description ''Valeriana rubra'' is a ...

. The

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

The caterpillars are coloured grey-brown, paler ventrally and have a pale dorsal line and clearly recognizable black point warts. The pupa is bright red. On the cremaster sit two short curved thorns.Larvae feed on a variety of plants, both wild and cultivated (see list below). This is one of the notorious cutworms and often severs or fatally damages plants at the base. The species overwinters as a full-grown larva in a chamber in the soil before

pupa A pupa (; : pupae) is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages th ...

ting in the spring. #''The flight season refers to the

British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...

. This may vary in other parts of the range.''

Recorded host plants

*''

Anemone ''Anemone'' () is a genus of flowering plants in the buttercup family Ranunculaceae. Plants of the genus are commonly called windflowers. They are native to the temperate and subtropical regions of all regions except Australia, New Zealand, and ...

'' *'' Artemisia'' *''Beta'' –

beet The beetroot (British English) or beet (North American English) is the taproot portion of a '' Beta vulgaris'' subsp. ''vulgaris'' plant in the Conditiva Group. The plant is a root vegetable also known as the table beet, garden beet, dinner ...

*''

Brassica rapa ''Brassica rapa'' is a plant species that has been widely cultivated into many forms, including the turnip (a root vegetable), komatsuna, napa cabbage, bomdong, bok choy, and rapini. ''Brassica rapa'' subsp. ''oleifera'' is an oilseed c ...

'' – turnip *''Fragaria'' –

strawberry The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown Hybrid (biology), hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit ...

*''Lactuca'' –

lettuce Lettuce (''Lactuca sativa'') is an annual plant of the family Asteraceae mostly grown as a leaf vegetable. The leaves are most often used raw in Green salad, green salads, although lettuce is also seen in other kinds of food, such as sandwiche ...

*''

Plantago ''Plantago'' is a genus of about 200 species of flowering plants in the family Plantaginaceae, commonly called plantains or fleaworts. The common name plantain is shared with the unrelated cooking plantain. Most are herbaceous plants, though a ...

'' - plantain *''

Polygonum ''Polygonum'' is a genus of about 130 species of flowering plants in the buckwheat and knotweed family Polygonaceae. Common names include knotweed and knotgrass (though the common names may refer more broadly to plants from Polygonaceae). In the ...

'' *''Quercus'' –

oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...

*''

Rubus ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. ...

'' – bramble *''

Rumex The docks and sorrels, genus ''Rumex'', are a genus of about 200 species of annual, biennial, and perennial herbs in the buckwheat family, Polygonaceae. Members of this genus are very common perennial herbs with a native almost worldwide distri ...

'' – docks, sorrels *''Solanum'' –

potato The potato () is a starchy tuberous vegetable native to the Americas that is consumed as a staple food in many parts of the world. Potatoes are underground stem tubers of the plant ''Solanum tuberosum'', a perennial in the nightshade famil ...

*''Spinacia'' –

spinach Spinach (''Spinacia oleracea'') is a leafy green flowering plant native to Central Asia, Central and Western Asia. It is of the order Caryophyllales, family Amaranthaceae, subfamily Chenopodioideae. Its leaves are a common vegetable consumed eit ...

*''

Stellaria ''Stellaria'' is a genus of about 190 species of flowering plants in the family Caryophyllaceae, with a cosmopolitan distribution. Common names include starwort, stitchwort and chickweed. Description ''Stellaria'' species are relatively small ...

'' *''Zea'' –

maize Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...

Subspecies

* ''A. e. corsica'' -

Corsica Corsica ( , , ; ; ) is an island in the Mediterranean Sea and one of the Regions of France, 18 regions of France. It is the List of islands in the Mediterranean#By area, fourth-largest island in the Mediterranean and lies southeast of the Metro ...

* ''A. e. exclamationis'' - Europe * ''A. e. informis'' Leech,

889 __NOTOC__ Year 889 ( DCCCLXXXIX) was a common year starting on Wednesday of the Julian calendar. Events By place Europe * Guy III, duke of Spoleto, defeats the Lombard king Berengar I at the Trebbia River, and is acclaimed as king ...

/small> - Japan Agrotis exclamationis-s.jpg, Profile Agrotis exclamationis-f.jpg, Front mark Agrotis exclamationis-o.jpg, Color variation

References

Chinery, Michael Michael Chinery (born 1938, in London) is an English naturalist. He studied in Cambridge where he graduated in natural sciences and anthropology. He edits '' Cecidology'', the journal of the British Plant Gall Society The British Plant Gall ...

(1986, reprinted 1991). ''Collins Guide to the Insects of Britain and Western Europe''. * Skinner, Bernard (1984). ''

The Colour Identification Guide to Moths of the British Isles ''The Colour Identification Guide to Moths of the British Isles (Macrolepidoptera)'' by Bernard Skinner is a single volume identification guide to the macromoths of Britain and Ireland. The first edition was published in 1984, and a second, revi ...

''.

External links

* *
''Lepiforum e.V.''
{{Taxonbar , from=Q771090 Agrotis Moths of Asia Moths of Europe Moths of Japan Moths of the Middle East Taxa named by Carl Linnaeus Moths described in 1758