''Agrimonia eupatoria'' is a species of

agrimony ''Agrimonia'' (from the Greek ), commonly known as agrimony, is a genus of 12–15 species of perennial plant, perennial herbaceous flowering plants in the family Rosaceae, native to the temperate regions of the Northern Hemisphere, with one sp ...

that is often referred to as common agrimony, church steeples or sticklewort. The whole

plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...

is dark green with numerous soft hairs. The soft hairs aid in the plant's seed pods sticking to any animal or person coming in contact with the plant. The flower spikes have a spicy odor like apricots. In the

language of flowers Floriography (language of flowers) is a means of cryptological communication through the use or arrangement of flowers. Meaning has been attributed to flowers for thousands of years, and some form of floriography has been practiced in tradition ...

, agrimony means thankfulness or gratitude. ''A. eupatoria'' is a foodplant for the

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...

s of the

snout moth The Pyralidae, commonly called pyralid moths, snout moths or grass moths, are a family (biology), family of Lepidoptera in the ditrysian Taxonomic rank, superfamily Pyraloidea. In many (particularly older) classifications, the grass moths (Cramb ...

Endotricha flammealis ''Endotricha flammealis'', the rose-flounced tabby, is a species of snout moth, family Pyralidae. Taxonomy The proposed subspecies ''carnealis'' and several supposed varieties seem to be indistinguishable from typical individuals found in Austri ...

''.

Description

Vegetative characteristics

The common agrimony grows as a

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...

herbaceous plant Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition o ...

and reached heights of up to . Its roots are deep

rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...

s, from which spring the stems. It is characterized by its typical serrated edged

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

Generative characteristics

The short-stemmed flowers appear from June to September, in long, spike-like,

racemose A raceme () or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the shoo ...

inflorescences In botany, an inflorescence is a group or cluster of flowers arranged on a plant's stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a main axis ( pe ...

. The single flower has an urn-shaped curved flower cup, the upper edge has several rows of soft, curved hook-shaped bristles, long. The

hermaphrodite A hermaphrodite () is a sexually reproducing organism that produces both male and female gametes. Animal species in which individuals are either male or female are gonochoric, which is the opposite of hermaphroditic. The individuals of many ...

flower has fivefold

radial symmetry Symmetry in biology refers to the symmetry observed in organisms, including plants, animals, fungi, and bacteria. External symmetry can be easily seen by just looking at an organism. For example, the face of a human being has a plane of symme ...

. There are five

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...

s present . There are five yellow, rounded

petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...

s. The petals and the five to 20

stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...

s rise above the tip of the flower cup . The two medium-sized

carpel Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more ...

s in the flower cups are sunk into, but not fused with it. The fruits are achenes approximately 0.6 cm (0.2 inch) in diameter and each have a number of hooks that enable it to cling to animal fur and clothing. Each achene may have one or two seeds. ''Agrimonia eupatoria'' is native to Europe and Southwestern Asia, where it grows in damp meadows, pasture, along stream banks, and among shrubs.

In folklore

Agrimony has been supposed to have magical, medical properties since the time of

Pliny the Elder Gaius Plinius Secundus (AD 23/24 79), known in English as Pliny the Elder ( ), was a Roman Empire, Roman author, Natural history, naturalist, and naval and army commander of the early Roman Empire, and a friend of the Roman emperor, emperor Vesp ...

. Common folklore held that it could cure

musket A musket is a muzzle-loaded long gun that appeared as a smoothbore weapon in the early 16th century, at first as a heavier variant of the arquebus, capable of penetrating plate armour. By the mid-16th century, this type of musket gradually dis ...

wounds by being brewed into ''"arquebusade water,"'' and ward off

witchcraft Witchcraft is the use of Magic (supernatural), magic by a person called a witch. Traditionally, "witchcraft" means the use of magic to inflict supernatural harm or misfortune on others, and this remains the most common and widespread meanin ...

. Traditional British folklore states that if a sprig of the plant was placed under a person's head, sleep would persist until it was removed.

Ecology

The flowers with their abundant pollen supply attract

hoverflies Hoverflies, also called flower flies or syrphids, make up the insect family (biology), family Syrphidae. As their common name suggests, they are often seen Hover (behaviour), hovering or nectaring at flowers; the adults of many species feed main ...

flies Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwin ...

and

honey bees A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to mainland Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the ...

. They also are an important food source for butterflies like the

grizzled skipper ''Pyrgus'' is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genera '' Burnsius'' and '' C ...

. The pollinated flowers develop fruits with burs. These attach to passing grazing animals, such as cattle, sheep and deer, and are dispersed over a large area. Agrimony is found usually in young

grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...

s, less than 50 years old. It is a wild host for a few insect pest species ('' Stigmella fragariella'' and ''Coroebus elatus'') that feed on loganberries, raspberries, and strawberries in Europe.

Phytochemistry

The plant contains

volatile oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...

flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...

apigenin Apigenin (4′,5,7-trihydroxyflavone), found in many plants, is a flavone compound that is the aglycone of several naturally occurring glycosides. It is a yellow crystalline solid that has been used to dye wool. Apigenin is abundant in parsl ...

luteolin Luteolin is a flavone, a type of flavonoid, with a yellow crystalline appearance. Luteolin is the main yellow dye from the '' Reseda luteola'' plant, used for dyeing since at least the first millennium B.C. Luteolin was first isolated in pure f ...

quercetin Quercetin is a plant flavonol from the flavonoid group of polyphenols. It is found in many fruits, vegetables, leaves, seeds, and grains; capers, red onions, and kale are common foods containing appreciable amounts of it. It has a bitter flavor ...

kaempferol Kaempferol (3,4′,5,7-tetrahydroxyflavone) is a natural flavonol, a type of flavonoid, found in a variety of plants and plant-derived foods including kale, beans, tea, spinach, and broccoli. It is also found in propolis extracts. Kaempferol i ...

, tiliroside,

triterpene Triterpenes are a class of terpenes composed of six isoprene units with the molecular formula C30H48; they may also be thought of as consisting of three terpene units. Animals, plants and fungi all produce triterpenes, including squalene, the pre ...

glycoside In chemistry, a glycoside is a molecule in which a sugar is bound to another functional group via a glycosidic bond. Glycosides play numerous important roles in living organisms. Many plants store chemicals in the form of inactive glycosides. ...

s, including euscaphic acid and tormentic acid,

phenolic acid Phenolic acids or phenolcarboxylic acids ? are phenolic compounds and types of aromatic acid compounds. Included in that class are substances containing a phenolic ring and an organic carboxylic acid function (C6-C1 skeleton). Two important nat ...

s, and 3%–21%

tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and Precipitation (chemistry), precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' is widel ...

Gallery

File:Agrimonia eupatoria (Agrimony) - Theodore Green - 26 1931 8.jpg, Image of ''Agrimony eupatoria'' by Theodore Green (1933)

Notes

References

* Howard, Michael. ''Traditional Folk Remedies'' (Century, 1987, pp 96–97)

External links

{{Taxonbar, from=Q156865 eupatoria Flora of temperate Asia Flora of North Africa Flora of Macaronesia Medicinal plants of Europe European witchcraft Plants described in 1753 Taxa named by Carl Linnaeus Herbs Superstitions of Great Britain