African Jacana
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The African jacana (''Actophilornis africanus'') is a wader in the family Jacanidae. It has long toes and long claws that enables it to walk on floating vegetation in shallow lakes, its preferred habitat. It is widely distributed in
sub-Saharan Africa Sub-Saharan Africa is the area and regions of the continent of Africa that lie south of the Sahara. These include Central Africa, East Africa, Southern Africa, and West Africa. Geopolitically, in addition to the list of sovereign states and ...
. For the origin and pronunciation of the name, see Jacanidae.


Taxonomy

The African jacanas was formally described in 1789 by the German naturalist
Johann Friedrich Gmelin Johann Friedrich Gmelin (8 August 1748 – 1 November 1804) was a German natural history, naturalist, chemist, botanist, entomologist, herpetologist, and malacologist. Education Johann Friedrich Gmelin was born as the eldest son of Philipp F ...
in his revised and expanded edition of
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
's ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
''. He placed it in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Parra'' and coined the binomial name ''Parra africana''. Gmelin based his description on that by the English ornithologist John Latham who in 1785 had described and illustrated the species in his ''A General Synopsis of Birds''. Gmelin and Latham gave the locality as "Africa": this was restricted to Ethiopia in 1915. The African jacana is now placed in the genus ''
Actophilornis ''Actophilornis'' is a genus of jacana. It contains two species restricted to Africa and its surrounding islands. Taxonomy The genus ''Actophilornis'' was introduced in 1925 by the American ornithologist Harry C. Oberholser with the African ...
'' that was introduced in 1925 by the American ornithologist Harry C. Oberholser. The genus name combines the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''aktē'' meaning "river bank" or "coastal strand", ''-philos'' meaning "-loving" and ''ornis'' meaning "bird". The species is considered to be
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised.


Description

The African jacanas is a conspicuous and unmistakable
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
. It measures in overall length. As in other jacanas, the female is on average larger than the male. Males can weigh from , averaging and females from , averaging . Alongside the similarly-sized Madagascar jacana, this appears to be the heaviest jacana species. They have chestnut upperparts with black wingtips, rear neck, and eyestripe. The underparts are also chestnut in the adults, only in juveniles they are white with a chestnut belly patch. The blue bill extends up as a coot-like head shield, and the legs and long toes are grey.


Behaviour


Food and feeding

African jacanas feed on
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s and other invertebrates picked from the floating vegetations or the surface of the water.


Breeding

African jacanas breed throughout sub-
Sahara The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
n
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
. It is sedentary apart from seasonal dispersion. It lays four black-marked brown eggs in a floating nest. The jacana has evolved a highly unusually polyandrous mating system, meaning that one female mates with multiple males and the male alone cares for the chicks. Such a system has evolved due to a combination of two factors: firstly, the lakes that the jacana lives on are so resource-rich that the relative energy expended by the female in producing each egg is effectively negligible. Secondly the jacana, as a bird, lays eggs, and eggs can be equally well incubated and cared for by a parent bird of either sex. This means that the rate-limiting factor of the jacana's breeding is the rate at which the males can raise and care for the chicks. Such a system of females forming harems of males is in direct contrast to the more usual system of leks seen in animals such as stags and grouse, where the males compete and display in order to gain harems of females. The parent that forms part of the harem is almost always the one that ends up caring for the offspring; in this case, each male jacana incubates and rears a nest of chicks. The male African jacana has therefore evolved some remarkable adaptations for parental care, such as the ability to pick up and carry chicks underneath its wings. African jacana (Actophilornis africanus) Kenya.jpg, Adult, Lake Baringo,
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
Actophilornis africana -Kakegawa Kacho-en, Kakegawa, Shizuoka, Japan -chick-8a.jpg, Chick at Kakegawa, Shizuoka,
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
African jacana (Actophilornis africanus) juvenile.jpg, Juvenile, Lake Baringo,
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
Actophilornis africanus MHNT.ZOO.2010.11.2.23.jpg, ''Actophilornis africanus'' - MHNT


References


External links

* African jacana
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q132201 African jacana African jacana African jacana African jacana African jacana Birds of East Africa