Afghan Snowfinch
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The Afghan snowfinch (''Pyrgilauda theresae'') or the Afghan ground-sparrow, bar-tailed snowfinch, Meinertzhagen's snowfinch, or Theresa's snowfinch, is a
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
bird of the sparrow family Passeridae,
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the northern parts of the
Hindu Kush The Hindu Kush is an mountain range in Central Asia, Central and South Asia to the west of the Himalayas. It stretches from central and eastern Afghanistan into northwestern Pakistan and far southeastern Tajikistan. The range forms the wester ...
mountains in Afghanistan. There are no major threats to the species despite its restricted range, so it is assessed as
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
. This species is mostly a seed-eater, supplementing its diet with some
insects Insects (from Latin ') are hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed ...
. It builds its nest in the burrows or hollows of ground-dwelling
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s, lined with hair or feathers.


Taxonomy and systematics

This species was first scientifically recorded relatively late, by
Richard Meinertzhagen Colonel Richard Meinertzhagen, CBE, DSO (3 March 1878 – 17 June 1967) was a British soldier, intelligence officer, and ornithologist. He had a decorated military career spanning Africa and the Middle East. He was credited with creating an ...
on a 1937 expedition with Salim Ali. Meinertzhagen formally described the species in '' Bulletin of the British Ornithologists' Club'' paper later that year, and gave it the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Montifringilla theresae''. He reported that he collected the
type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
of the species in the Shibar Pass, between
Bamyan Bamyan (), also spelled Bamian or Bamiyan, is the capital of Bamyan Province in central Afghanistan. Its population of approximately 100,000 people makes it the largest city in Hazarajat. Bamyan is at an altitude of about above sea level. The ...
and
Kabul Kabul is the capital and largest city of Afghanistan. Located in the eastern half of the country, it is also a municipality, forming part of the Kabul Province. The city is divided for administration into #Districts, 22 municipal districts. A ...
. The name he gave the species was after his cousin and companion, Theresa Clay, an expert on bird lice. While Meinertzhagen described several dozen species and
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
(including others named after Clay), he was later found to have stolen specimens and falsified records, and this may be the only verifiably genuine taxon he described. Meinertzhagen placed this species in the genus '' Montifringilla'', which some classifications continue to place it in, along with about eight species of 'snowfinches'. Among these species, it is most similar to Blanford's snowfinch, which has similar plumage and essentially the same nesting habits. Some authors split this species, Blanford's snowfinch, and the other snowfinches of southern and central Asia as the genus ''Pyrgilauda'', on the basis that their morphological traits, as well as habitat and ecology, are substantially different. (This is an arbitrary distinction, as there can be little doubt that ''Montifringilla'' in the broad sense is
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
.) One of the more prominent works to follow this classification recently, the ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
'', called the species of their genus ''Pyrgilauda'' 'ground-sparrows', on account of their different habitat and behaviour from the northern 'snowfinches'. No subspecies have been described.


Description

It is long, weighing . The wingspan ranges from , and bills measure from . Females are slightly smaller on average than males. The male is grey-brown with some white in the wings and a black face-mask and two-pronged patch on the throat. The female is a buff-tinged brown, with a weaker, greyer face mask and less white in the wings. There are short dark streaks on the mantle and a white subterminal band on the tail feathers other than the central pair. There is white on the upperwing coverts, secondaries, and inner primaries. The male has a brick red iris. The plumage of juvenile birds after fledging has not been described, but can be assumed to be similar to the female's plumage. The only similar species that occur in the Afghan snowfinch's range are the white-winged snowfinch and the desert finch. It can be distinguished from the former by the smaller white patches on the wings and an overall more brownish plumage. While it is similar in general appearance to the latter species, the Afghan snowfinch is more streaked, has stronger facial markings, and has a smaller bill, among other differences. The flight of the Afghan snowfinch is heavy and straight. The alarm call is a sharp ', and they make soft ' calls in flight, and a stridulant '.


Distribution and habitat

The Afghan snowfinch is the only species of bird known to be endemic to Afghanistan. It is found only in some northern parts of the
Hindu Kush The Hindu Kush is an mountain range in Central Asia, Central and South Asia to the west of the Himalayas. It stretches from central and eastern Afghanistan into northwestern Pakistan and far southeastern Tajikistan. The range forms the wester ...
mountains, where it occurs at elevations of . Besides the Shibar Pass, it is known from Deh Sabz and Unai Pass, and a few other localities between 67° and 69° E in the northerly ranges of the Hindu Kush. This species disperses in the winter especially after heavy snowfalls, and moves slightly beyond its breeding range, into lower altitudes and northwards into
Badghis Province Bādghīs () is one of the thirty-four provinces of Afghanistan, located in the northwest of the country, on the border with Turkmenistan. It is considered to be one of the country's most underdeveloped provinces, with the highest poverty rate. T ...
. It has been recorded on occasion as a
vagrant Vagrancy is the condition of wandering homelessness without regular employment or income. Vagrants usually live in poverty and support themselves by travelling while engaging in begging, scavenging, or petty theft. In Western countries, ...
in southern
Turkmenistan Turkmenistan is a landlocked country in Central Asia bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the south and southwest and the Caspian Sea to the west. Ash ...
. Its habitats are stony mountain slopes, plateaux, and open hillsides in the passes. Despite having a relatively small range and population, it is not thought to have an unstable population or significant threats, so it is assessed as a species of
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
. This species is among those protected by Band-e Amir National Park, Afghanistan's first national park, which encompasses a large area of the Hindu Kush near Bamyan.


Behaviour and ecology

In winter, the Afghan snowfinch forms large flocks of dozens or hundreds, sometimes mixed with snowfinches of other species, rock sparrows, and various
lark Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occ ...
s. This species feeds mostly on small seeds, from plants such as '' Carex pachystylis'', '' Convolvulus divaricatus'', and '' Thuspeinantha persica'', and it will also eat insects such as
ant Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...
s and weevils. The Afghan snowfinch builds
nest A nest is a structure built for certain animals to hold Egg (biology), eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of ...
s in the burrows and hollows made by rodents, including
ground squirrel Ground squirrels are rodents of the squirrel family (Sciuridae) that generally live on the ground or in burrows, rather than in trees like the tree squirrels. The term is most often used for the medium-sized ground squirrels, as the larger ones ar ...
s,
marmot Marmots are large ground squirrels in the genus ''Marmota'', with 15 species living in Asia, Europe, and North America. These herbivores are active during the summer, when they can often be found in groups, but are not seen during the winter, w ...
s, and in one recorded case Williams' jerboa (''Allactaga williamsi''). In particular, it is associated with the yellow ground squirrel (''Spermophilus fulvus''). Several of the other southern snowfinch species have similar nesting preferences, as does at least one species that occurs in the Afghan snowfinch's range, the isabelline wheatear. The materials used to line the burrows it nests in are the hairs of squirrels,
sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...
, and dromedaries; and
feather Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and an exa ...
s. Nests are built at the far end of the burrows, as protection from predators. The young hatch blind and helpless, with exposed pink skin and only a few, light-coloured tufts of down. Young give a soft call in response to adults calling at the entrance of a nest.


References


Works cited

* * *


External links

* {{Taxonbar, from=Q27075723 Birds described in 1937 Birds of Afghanistan Endemic fauna of Afghanistan Pyrgilauda