Aetosaurus
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''Aetosaurus'' is an extinct
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
pseudosuchian Pseudosuchia, from Ancient Greek ψεύδος (''pseúdos)'', meaning "false", and σούχος (''soúkhos''), meaning "crocodile" is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely relat ...
reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
belonging to the
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Aetosauria. It is generally considered to be the most primitive aetosaur. Three species are currently recognized: ''A. ferratus'', the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
from
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
and
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
; ''A. crassicauda'' from Germany; and ''A. arcuatus'' from eastern
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. Additional specimens referred to ''Aetosaurus'' have been found in the Chinle Group of the southwestern
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
, and the
Fleming Fjord Formation The Fleming Fjord Formation, alternatively called the Fleming Fjord Group is an Upper Triassic geological formation in the northeastern coast of Jameson Land, Greenland. It consists of terrestrial sediments and is known for its fossil content. ...
of
Greenland Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...
. Specimens of ''Aetosaurus'' occur in
Norian The Norian is a division of the Triassic geological period, Period. It has the rank of an age (geology), age (geochronology) or stage (stratigraphy), stage (chronostratigraphy). It lasted from ~227.3 to Mya (unit), million years ago. It was prec ...
-age strata.


Description

''Aetosaurus'' was a small, primitive aetosaur. Unlike more derived aetosaurs such as ''
Desmatosuchus ''Desmatosuchus'' (, from Greek language, Greek δεσμός ''desmos'' 'link' + σοῦχος ''soûkhos'' 'crocodile') is an extinct genus of archosaur belonging to the Order Aetosauria. It lived during the Late Triassic. Description ''Desma ...
'' or '' Typothorax'', the
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
was long and narrow and lacked spikes. The paramedian
scute A scute () or scutum (Latin: ''scutum''; plural: ''scuta'' "Scutum (shield), shield") is a bony external plate or scale overlaid with horn, as on the shell of a turtle, the skin of crocodilians, and the feet of Bird anatomy#Scales, birds. The ter ...
s that covered the back (with one row on each side of the vertebrae) are considerably wider than they are long. The lateral scutes, which are beneath the paramedian and formed a row on either side of the animal, do not bear any spikes or other projections.


Behaviour

A 2023 study based on a fossil assemblage found in Kaltental, suggested that juvenile ''A. ferratus'' were likely
gregarious Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother was ...
animals, possibly to increase their chances of survival and to deter predators.


Species

''Aetosaurus'' was first named, with the description of the type species ''A. ferratus'', in 1877 by German
paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
Oscar Fraas. At the time, ''Aetosaurus'' was known from 22 articulated skeletons that had been found in the Lower Stubensandtein of Germany. Thirty years later, Fraas' son
Eberhard Eberhard is an old Germanic name meaning the strength or courage of a wild boar. People First name *Eberhard of Friuli (815–866), Duke and key figure in the Carolingian Empire *Eberhard of Béthune (died 1212), Flemish grammarian *Eberhard I, Du ...
described a second species, ''A. crassicauda'', also from Germany. ''A. crassicauda'' can be distinguished from ''A. ferratus'' by its larger size; ''A. crassicauda'' reached a maximum length of while ''A. ferratus'' reached a length of up to . In addition to the Stubensandtein in Germany, ''A. ferratus'' is also known from the Calcare di Zorzino Formation in Cene, Italy. Specimens of ''Aetosaurus'' that have been recovered from the Fleming Fjord Formation in Greenland likely represent ''A. ferratus''. Some material from the Chinle Group in the southwestern United States probably represent ''A. ferratus'', as well. In 1998, the genus ''Stegomus'' was synonymized with ''Aetosaurus''. In 1896, paleontologist
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of paleontology. A prolific fossil collector, Marsh was one of the preeminent paleontologists of the nineteenth century. Among his legacies are the discovery or ...
named ''Stegomus arcuatus'' from a
cast Cast may refer to: Music * Cast (band), an English alternative rock band * Cast (Mexican band), a progressive Mexican rock band * The Cast, a Scottish musical duo: Mairi Campbell and Dave Francis * ''Cast'', a 2012 album by Trespassers William ...
of an aetosaur known as YPM 1647 from the New Haven Formation in the Fair Haven Heights neighborhood of New Haven. This cast consisted of the dorsal carapace. Several other casts preserving the surface of the skull and tail have been found in the Passaic Formation of Hunterdon and Somerset counties,
New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
and an outcrop of the Lower Sanford Formation at the Triangle Brick Co. Quarry in
Durham County, North Carolina Durham County is a county located in the U.S. state of North Carolina. As of the 2020 census, the population was 324,833, making it the sixth-most populous county in North Carolina. Its county seat is Durham, which is the only incorporate ...
. ''Stegomus arcuatus'' was found to be synonymous with ''Aetosaurus'' on the basis of several similarities, including a lack of spikes and a distinctive radial pattern of grooves on some of the caudal scutes. ''A. arcuatus'' has paramedian scutes that are much wider than they are long, even in comparison to other species of ''Aetosaurus''. There is very little pitting on the surface of the scutes, although the porosity of the sandstone that makes up the casts has been mistaken for pitting. The tail narrows significantly past the base. The carapace is "waisted", meaning that it narrows in front of the
pelvis The pelvis (: pelves or pelvises) is the lower part of an Anatomy, anatomical Trunk (anatomy), trunk, between the human abdomen, abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also c ...
. The following
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
is simplified after an analysis presented by Julia B. Desojo, Martin D. Ezcurra and Edio E. Kischlat (2012).


References


External links


''Aetosaurus''
in the
Paleobiology Database The Paleobiology Database (PBDB) is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database originated in the NCEAS-funded Phanerozoic Marine Pale ...
{{Taxonbar, from=Q3496825 Late Triassic reptiles of Europe Aetosauria Late Triassic reptiles of Africa Aetosaurs of North America Late Triassic pseudosuchians Prehistoric pseudosuchian genera Fossil taxa described in 1877 Fossil taxa described in 1887