Aelurillus Basseleti
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''Aelurillus basseleti'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
jumping spider Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. , this family contained over 600 species description, described genus, genera and over 6,000 described species, making it the largest family of spide ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Aelurillus ''Aelurillus'' is a genus of spiders in the family Salticidae (jumping spiders). Description Species of the genus ''Aelurillus'' are typically about 7 mm long in females, and up to five mm in males. They are stout, squat-shaped and rather f ...
'' that has been found in Algeria, Morocco and Tunisia. Originally named ''Salticus basseleti'', the spider was first identified in 1846 by
Hippolyte Lucas Pierre-Hippolyte Lucas (17 January 1814 – 5 July 1899) was a French entomologist. Lucas was an assistant-naturalist at the Muséum national d'Histoire naturelle. From 1839 to 1842 he studied fauna as part of the scientific commission on the ex ...
, but the original male
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
has been lost. The female was first described in 2006. The spider is small and hard to distinguish from the related species '' Aelurillus luctuosus'' and '' Aelurillus monardi''. The dark brown
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
is typically between long and the grey-yellow
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
is between long, the female being larger than the male. The carapace has a single stripe down the middle. While the female is hard to distinguish compared to others in the genus, the male spider has distinctive white or tawny bands on the
clypeus The clypeus is one of the sclerites that make up the face of an arthropod. In insects, the clypeus delimits the lower margin of the face, with the labrum articulated along the ventral margin of the clypeus. The mandibles bracket the labrum, but ...
. The male has a curved
embolus An embolus (; : emboli; from the Greek ἔμβολος "wedge", "plug") is an unattached mass that travels through the circulatory system, bloodstream and is capable of creating blockages. When an embolus Vascular occlusion, occludes a blood vess ...
that is sufficiently varied between individual spiders that it is not sufficiently specific to identify the species.


Taxonomy

''Aelurillus basseleti'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
jumping spider Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. , this family contained over 600 species description, described genus, genera and over 6,000 described species, making it the largest family of spide ...
, a member of the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Salticidae Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. , this family contained over 600 species description, described genus, genera and over 6,000 described species, making it the largest family of spide ...
, that was first described by the French
arachnologist Arachnology is the scientific study of arachnids, which comprise spiders and related invertebrates such as scorpions, pseudoscorpions, harvestmen, ticks, and mites. Those who study spiders and other arachnids are arachnologists. More narrowly ...
Hippolyte Lucas Pierre-Hippolyte Lucas (17 January 1814 – 5 July 1899) was a French entomologist. Lucas was an assistant-naturalist at the Muséum national d'Histoire naturelle. From 1839 to 1842 he studied fauna as part of the scientific commission on the ex ...
in 1846. It was initially named ''Salticus basseleti'' and allocated to the genus ''
Salticus ''Salticus'' (from Latin “''saltus''” – leap or jump) is a genus of the family Salticidae (the jumping spiders). ''Salticus'' is the type genus for the family Salticidae. Description Coloration is determined by various scales (modified se ...
''. The genus name was derived from the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
word saltus, meaning something that jumps. In 1876,
Eugène Simon Eugène Louis Simon (; 30 April 1848 – 17 November 1924) was a French naturalist who worked particularly on insects and spiders, but also on birds and plants. He is by far the most prolific spider Taxonomy (biology), taxonomist in history, ...
moved the species to the genus ''Aelurops'' and then, in 1880, Pietro Pavesi moved it to the genus ''Ictidops'' before Simon finally moved it to ''
Aelurillus ''Aelurillus'' is a genus of spiders in the family Salticidae (jumping spiders). Description Species of the genus ''Aelurillus'' are typically about 7 mm long in females, and up to five mm in males. They are stout, squat-shaped and rather f ...
'' in 1886. The genus ''Aelurillus'' was first described by
Eugène Simon Eugène Louis Simon (; 30 April 1848 – 17 November 1924) was a French naturalist who worked particularly on insects and spiders, but also on birds and plants. He is by far the most prolific spider Taxonomy (biology), taxonomist in history, ...
one year before, in 1885. The genus name derives from the Greek word for cat. It was placed in the subtribe Aelurillina in the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Aelurillini, both named after the genus, by
Wayne Maddison Wayne Paul Maddison (born 1958) is a Canadian evolutionary biologist, arachnologist, and biological illustrator. He is Canada Research Chair in Biodiversity and a professor at the departments of zoology and botany at the University of British ...
in 2015. These were allocated to the
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
Saltafresia. In 2017, the genus was grouped with nine other genera of jumping spiders under the name Aelurillines by
Jerzy Prószyński Jerzy Prószyński (born 1935 in Warsaw) is a Polish arachnologist specializing in systematics of jumping spiders (family Salticidae). He is a graduate of the University of Warsaw, a long-term employee of the Siedlce University of Natural Sc ...
.


Description

Initially, only the male was described. The spider is small, about long. The male has a dark brown
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
that is typically long and wide. Two white stripes cross the otherwise black eye field and extend along the rest of the carapace. Its
clypeus The clypeus is one of the sclerites that make up the face of an arthropod. In insects, the clypeus delimits the lower margin of the face, with the labrum articulated along the ventral margin of the clypeus. The mandibles bracket the labrum, but ...
is brown and covered with dense hairs that are either white or tawny. The hairs form wide bands which are distinctive for the species. Its
chelicerae The chelicerae () are the arthropod mouthparts, mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated ...
are brown with short brown hairs. The spider's
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
is long and wide. It is grey-yellow on the top and has a brown underside with a wide white band across the middle. Its
spinneret A spinneret is a silk-spinning organ of a spider or the larva of an insect. Some adult insects also have spinnerets, such as those borne on the forelegs of Embioptera. Spinnerets are usually on the underside of a spider's opisthosoma, and ar ...
s are grey-brown and its
legs A leg is a weight-bearing and locomotive anatomical structure, usually having a columnar shape. During locomotion, legs function as "extensible struts". The combination of movements at all joints can be modeled as a single, linear element cap ...
are yellow. Its
pedipalp Pedipalps (commonly shortened to palps or palpi) are the secondary pair of forward appendages among Chelicerata, chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to ...
s are also yellow and have a covering of white hairs. The
palpal bulb The two palpal bulbs – also known as palpal organs and genital bulbs – are the copulatory organs of a male spider. They are borne on the last segment of the pedipalps (the front "limbs" of a spider), giving the spider an appearance often descr ...
has two short tibial apophyses and distinctive blended appendages that extend from the middle. The
tegulum The two palpal bulbs – also known as palpal organs and genital bulbs – are the copulatory organs of a male spider. They are borne on the last segment of the pedipalps (the front "limbs" of a spider), giving the spider an appearance often desc ...
is rounded with a long coiled
embolus An embolus (; : emboli; from the Greek ἔμβολος "wedge", "plug") is an unattached mass that travels through the circulatory system, bloodstream and is capable of creating blockages. When an embolus Vascular occlusion, occludes a blood vess ...
which has a hooked end. The shape of the embolus shows sufficient variation that it is not a distinguishing feature for the species. The female was not described until 2006. It is large, with a carapace that is long and wide. It is similar in colour but has a covering of white and yellow scales that can form a pattern reminiscent of a net. The abdomen is also larger, with a length of and a width of . It is similar in colour on the top, but the underside has a similar pattern of brown and white scales. Although the clypeus is similar in colour, much of the remaining features are darker. The chelicerae are darker brown, the spinnerets are brown-grey and the legs have dark brown patches on an otherwise brown background. Its
epigyne The epigyne or epigynum is the external genital structure of female spiders. As the epigyne varies greatly in form in different species, even in closely related ones, it often provides the most distinctive characteristic for recognizing species. ...
has a pocket and short copulatory ducts. The spider is typical of the genus and can be confused with other related species. It is superficially similar, particularly in its patterning. It can be distinguished from the related '' Aelurillus luctuosus'' and '' Aelurillus monardi'' by the band of hairs on the clypeus. The females are almost indistinguishable. Similarly, there is too large a variation in the male palpal bulb to enable clear differentiation.


Distribution

''Aelurillus'' spiders have a wide
distribution Distribution may refer to: Mathematics *Distribution (mathematics), generalized functions used to formulate solutions of partial differential equations *Probability distribution, the probability of a particular value or value range of a varia ...
, mainly in the
Palearctic realm The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. The ...
, with 34 living in the area around the
Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...
. ''Aelurillus basseleti'' has been found in Algeria, Morocco and Tunisia. The spider was first identified by Lucas in
Constantine, Algeria Constantine (), also spelled Qacentina or Kasantina, is the capital of Constantine Province in northeastern Algeria. During Roman times it was called Cirta and was renamed "Constantina" in honour of Emperor Constantine the Great. Located somewh ...
. This
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
was subsequently lost. Examples have also been found in locations around the country including
Zemmouri Zemmouri is a town and commune in the Bordj Menaïel District of Boumerdès Province, Algeria. As of 2008, the population of the municipality is 26,408. Villages The villages of the commune of ''Zemmouri'' are: History The coastal site of Zem ...
in 1984,
El Harrach El Harrach (Berber: ⵍⵃⴻⵔⵔⴰⵛ , formerly ''Maison-Carrée'') is a suburb of the Algerian capital Algiers. The town is home to USM El Harrach Union Sportive de la Médina d'El Harrach (), known as USM El Harrach or simply USMH for ...
in 1985,
Reghaïa Reghaïa is a municipality in Algiers Province, Algeria. It is located in Rouïba District, Rouïba districts of Algeria, district and is the most populated municipality of the district. Geography It is located in the northern part of the plain ...
in 1988 and
Taghit Taghit () is a town and commune in Taghit District, Béchar Province, in western Algeria. The town is an oasis watered by the underground Oued Zousfana, which runs along beside the dunes of the Grand Erg Occidental. According to the 2008 cens ...
in 1989. The spider has been found in Tunisia, first noted in
Aïn Draham Aïn Draham ( ') is a city in northwestern Tunisia in the Jendouba Governorate, situated 25 kilometers south of Tabarka near the border with Algeria. Historically a military outpost, summer resort and souq, today it is a regional economic hub. ...
by Eugène Simon in 1885. It has been identified in
Jendouba Jendouba ( ; Formerly known as Souk El Arba until 30 April 1966) is a city in northwestern Tunisia, and capital of the Jendouba Governorate. It is an important crossroads with many road links to other towns such as El Kef, Tabarka, Ain Draham ...
,
Nabeul Nabeul (; ; Tamazight: ⵏⴰⴱⴻⵍ) is a coastal town located in northeastern Tunisia, on the south coast of the Cape Bon peninsula and surrounded by the Mediterranean Sea on both sides. It is the first seaside resort in Tunisia. It is know ...
and
Tunis Tunis (, ') is the capital city, capital and largest city of Tunisia. The greater metropolitan area of Tunis, often referred to as "Grand Tunis", has about 2,700,000 inhabitants. , it is the third-largest city in the Maghreb region (after Casabl ...
. In 2003, Jerzy Prószyński described a male spider from the
Levant The Levant ( ) is the subregion that borders the Eastern Mediterranean, Eastern Mediterranean sea to the west, and forms the core of West Asia and the political term, Middle East, ''Middle East''. In its narrowest sense, which is in use toda ...
, which he saw as another example of the species. However, when this example was re-examined by Galina Azarkina and Dmitri Logunov in 2006, it was found to be the related ''Aelurillus luctuosus''. In 2013, Liviu Moscaliuc found the first example in Morocco, near Sidi Kaouki. In terms of
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
, the spider inhabits a wide range of environments. Examples have been found in open grassland amongst flowering plants like ''
Oxalis pes-caprae ''Oxalis pes-caprae'', commonly known as African wood-sorrel, Bermuda buttercup, Bermuda sorrel, buttercup oxalis, Cape sorrel, English weed, goat's-foot, sourgrass, soursob or soursop; ; (حميضة), is a species of tristylous yellow-floweri ...
'', in areas of shrubs like ''
Cistus salviifolius ''Cistus salviifolius'', common names sage-leaved rock-rose, salvia cistus or Gallipoli rose, is a shrub of the family Cistaceae. Etymology The genus name ''Cistus'' derives from the Ancient Greek words ''κίσθος'' (''kisthos'') meaning bas ...
'', near to pools found in
dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, flat ...
s, in
palm Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae ** List of Arecaceae genera **Palm oil * Several other plants known as "palm" Palm or Palms may also refer to: Music ...
groves and within plantations of
cedar Cedar may refer to: Trees and plants *''Cedrus'', common English name cedar, an Old-World genus of coniferous trees in the plant family Pinaceae * Cedar (plant), a list of trees and plants known as cedar Places United States * Cedar, Arizona ...
trees.


References


Citations


Bibliography

* * * * * * * * * * {{Taxonbar, from=Q2255476 Spiders of Algeria Spiders of Morocco Salticidae Spiders described in 1846 Taxa named by Hippolyte Lucas