Adjutant Stork
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''Leptoptilos'' is a genus of very large tropical
stork Storks are large, long-legged, long-necked wading birds with long, stout bills. They belong to the family Ciconiidae, and make up the order Ciconiiformes . Ciconiiformes previously included a number of other families, such as herons and ibise ...
s, commonly known as adjutants. The name means thin (''lepto'') feather (''ptilos''). Two species are resident breeders in southern
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, and the
marabou stork The marabou stork (''Leptoptilos crumenifer'') is a large wading bird in the stork family Ciconiidae native to sub-Saharan Africa. It breeds in both wet and arid habitats, often near human habitation, especially landfill sites. It is sometim ...
is found in
Sub-Saharan Africa Sub-Saharan Africa is the area and regions of the continent of Africa that lie south of the Sahara. These include Central Africa, East Africa, Southern Africa, and West Africa. Geopolitically, in addition to the list of sovereign states and ...
. These are huge
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s, typically 110–150 cm tall with a 210–250 cm wingspan. The three species each have a black upper body and wings, and white belly and undertail. The head and neck are bare like those of a
vulture A vulture is a bird of prey that scavenges on carrion. There are 23 extant species of vulture (including condors). Old World vultures include 16 living species native to Europe, Africa, and Asia; New World vultures are restricted to Nort ...
. The huge
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Pl ...
is long and thick. Juveniles are a duller, browner version of the adult. ''Leptoptilos'' storks are gregarious colonial breeders in
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
s, building large stick nests in trees. They feed on
frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
s,
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s, young birds,
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s and
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s. They are frequent
scavenger Scavengers are animals that consume Corpse decomposition, dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a he ...
s, and the naked head and neck are adaptations to this, as are those of the vultures with which they often feed. A feathered head would become rapidly clotted with blood and other substances when a scavenging bird's head was inside a large corpse, and the bare head is easier to keep clean. Most storks fly with neck outstretched, but the three ''Leptoptilos'' storks retract their necks in flight like a
heron Herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 75 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genus ''Botaurus'' are referred to as bi ...
.


Taxonomy and species

The genus ''Leptoptilos'' was introduced in 1831 by the French naturalist
René Lesson René Primevère Lesson (20 March 1794 – 28 April 1849) was a French surgery, surgeon, natural history, naturalist, ornithologist, and herpetologist. Biography Lesson was born at Rochefort, Charente-Maritime, Rochefort, and entered the Naval ...
. The genus name combines the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''leptos'' meaning "delicate" or "slender" with ''ptilon'' meaning "feather". The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
was subsequently designated as the
greater adjutant The greater adjutant (''Leptoptilos dubius'') is a member of the stork family, Ciconiidae. Its genus includes the lesser adjutant of Asia and the marabou stork of Africa. Once found widely across southern Asia and mainland southeast Asia, t ...
by
George Robert Gray George Robert Gray (8 July 1808 – 6 May 1872) was an English zoology, zoologist and author, and head of the Ornithology, ornithological section of the British Museum, now the Natural History Museum, London, Natural History Museum, London f ...
. The genus contains three extant species.


Fossils

There is an ample
fossil record A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
of this genus. Many fossils members of the genus were much larger than living species, standing as tall as a man, with the earliest being ''
Leptoptilos falconeri ''Leptoptilos falconeri'' is an extinct species of large-bodied stork that existed during the Pliocene, having persisted until just over 2.58 million years ago. Although not the oldest fossil species of the genus ''Leptoptilos'' (as several date ...
'' from the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
on the Southeast Asian islands of Java (''L''. ''titan'') and Flores ( ''L''. ''robustus''). *†''
Leptoptilos falconeri ''Leptoptilos falconeri'' is an extinct species of large-bodied stork that existed during the Pliocene, having persisted until just over 2.58 million years ago. Although not the oldest fossil species of the genus ''Leptoptilos'' (as several date ...
'' (Early to Late Pliocene of south Asia and east Africa) *†'' Leptoptilos indicus'' (Late Pliocene of
Siwalik The Sivalik Hills, also known as Churia Hills, are a mountain range of the outer Himalayas. The literal translation of "Sivalik" is 'tresses of Shiva'. The hills are known for their numerous fossils, and are also home to the Soanian Middle Pale ...
, India) – formerly ''Cryptociconia indica'', may be the same as ''L. falconeri'' (Louchart ''et al.'' 2005) *†'' Leptoptilos lui'' (
Middle Pleistocene The Chibanian, more widely known as the Middle Pleistocene (its previous informal name), is an Age (geology), age in the international geologic timescale or a Stage (stratigraphy), stage in chronostratigraphy, being a division of the Pleistocen ...
of
Jinniushan Jinniushan () is a Middle Pleistocene paleoanthropological site, dating to around 260,000 BP, most famous for its archaic hominin fossils. The site is located near Yingkou, Liaoning, China. Several new species of extinct birds were also discovere ...
, Liaoning, China) *†''
Leptoptilos patagonicus ''Leptoptilos'' is a genus of very large tropical storks, commonly known as adjutants. The name means thin (''lepto'') feather (''ptilos''). Two species are resident breeders in southern Asia, and the marabou stork is found in Sub-Saharan Afric ...
'' (Puerto Madryn
Late Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million ye ...
of
Valdés Peninsula The Valdes Peninsula (Spanish: ''Península Valdés'') is a peninsula into the Atlantic Ocean in the Viedma Department of north-east Chubut Province, Argentina. It is an important nature reserve which was listed as a World Heritage Site by UNESC ...
, Argentina) *†'' Leptoptilos pliocenicus'' (Early Pliocene of Odesa, Ukraine and Urugus, Ethiopia to Late Pliocene of
Koro Toro Koro Toro is a settlement in the southern Borkou Region of Chad. It hosts the Koro Toro Airport and a "notorious" maximum security desert prison used by the Chadian government to detain captured fighters of Boko Haram and Chadian rebel groups. Acco ...
, Chad and Olduvai, Tanzania) – includes ''L.'' cf. ''falconeri'', may be the same as ''L. falconeri'' *†'' Leptoptilos richae'' (Beglia Late Miocene of Bled ed Douarah, Tunisia, and
Wadi Moghara Wadi ( ; ) is a river valley or a wet (ephemeral) riverbed that contains water only when heavy rain occurs. Wadis are located on gently sloping, nearly flat parts of deserts; commonly they begin on the distal portions of alluvial fans and ext ...
, Egypt?) *†''
Leptoptilos robustus ''Leptoptilos robustus'' (from reek: thin, slender+ reek: soft featherand atin: strong is an extinct species of large-bodied stork belonging to the genus ''Leptoptilos'' that lived on the island of Flores in Indonesia during the Pleistocen ...
'' (Pleistocene,
Flores Flores is one of the Lesser Sunda Islands, a group of islands in the eastern half of Indonesia. Administratively, it forms the largest island in the East Nusa Tenggara Province. The area is 14,250 km2. Including Komodo and Rinca islands ...
, Indonesia) *†''
Leptoptilos titan ''Leptoptilos'' is a genus of very large tropical storks, commonly known as adjutants. The name means thin (''lepto'') feather (''ptilos''). Two species are resident breeders in southern Asia, and the marabou stork is found in Sub-Saharan Afric ...
'' (Notopuro Middle/Late Pleistocene of Watualang, Java, Indonesia) *†''Leptoptilos'' sp. (
Ngorora Formation The Ngorora Formation is a geological formation in Kenya preserving fossils dating to the Miocene. The uppermost member of the formation shows sign of a faunal turnover that occurred around 11 to 10.5 million years ago, coinciding with faunal ch ...
Late Miocene of
Baringo District Baringo District was an administrative district in the Rift Valley Province of Kenya. Its capital town was Kabarnet. The district had a population of 264,978 (1999 census) and an area of . The district was created by the colonial government. In 2 ...
, Kenya: Louchart ''et al.'' 2005) †''
Leptoptilos siwalicensis ''Leptoptilos'' is a genus of very large tropical storks, commonly known as adjutants. The name means thin (''lepto'') feather (''ptilos''). Two species are resident breeders in southern Asia, and the marabou stork is found in Sub-Saharan Afric ...
'' from the
Siwalik The Sivalik Hills, also known as Churia Hills, are a mountain range of the outer Himalayas. The literal translation of "Sivalik" is 'tresses of Shiva'. The hills are known for their numerous fossils, and are also home to the Soanian Middle Pale ...
deposits (Late Miocene? to Late Pliocene) may belong to this genus or to a closely related one (Louchart ''et al.'' 2005).


References

{{Authority control Bird genera Taxa described in 1831 Taxa named by René Lesson