Adeilosmilus

''Amphimachairodus'' is an extinct genus of large machairodonts. It is also a member of the tribe Homotherini within Machairodontinae and is most closely related to such species as '' Xenosmilus'', ''Homotherium'' itself, and '' Nimravides''. It inhabited Eurasia, Northern Africa and North America during the late Miocene epoch.

Description

There was marked sexual dimorphism in ''A. giganteus'', with males being much larger than females.

The species ''Amphimachairodus coloradensis'', from the United States (formerly ''Machairodus coloradensis'') was a significantly large animal, about at the shoulder, according to skeletal and life reconstructions, potentially making it one of the largest known felids. All ''Amphimachairodus'' species have a developed mandibular flange, however, ''A. colaradensis'' is distinguishable from ''A. giganteus'' and ''A. kurteni'' by subtle differences in the shape of the mandible and placement of lower carnassials.
In size and proportions, the Eurasian species ''A. giganteus'' was remarkably similar to a modern lion or tiger and had a shoulder height of . This species has a skull length of around . The African species ''A. kabir'' (formerly ''Machairodus kabir'', from Arabic ''kabir'' = "big") is suggested to have weighed over . This would make it comparable in size to '' Xenosmilus'', '' Machairodus horribillis'' and slightly smaller than ''Smilodon populator''. In 2022, this species was proposed to be reassigned to a separate genus, called ''Adeilosmilus''.

''Amphimachairodus'' was about long and probably hunted as an ambush predator. Its legs were too short to sustain a long chase, but it most likely was a good jumper. It probably used its canines to cut open the throat of its prey, severing the major arteries and possibly crushing the windpipe. Its teeth were rooted to its mouth and were not as delicate as those of most other saber-toothed cats of the time, which had extremely long canines that hung out of their mouths. The fangs of ''Amphimachairodus'', however, were able to easily fit in its mouth comfortably while being long enough to be effective for hunting. ''Amphimachairodus'' also possessed a longer tail than most other large machairodont genera.

Skull

This specimen was from a large male ''A. giganteus'' with the skull measuring from the Late Miocene in China, comparable to a male lion or tiger. Deformation of the skull through natural fossilization processes has changed the shape slightly, making it asymmetrical, but overall it remains an excellent specimen for studying the cranial morphology of this particular genus and species.

For felines, this skull is rather long, but rivaled by the skulls of the two largest species of extant cats: the lion and tiger. When compared with the skull of a regular lion, it is long and very narrow, particularly in the muzzle and width of the zygomatic arches. Its sagittal crest is well pronounced. Compared with other machairodonts, the canines are stout and capable of large amounts of stress. This characteristic is slightly remodeled in females, whose canines are slimmer and generally longer. Compared with females, the orbit of males are smaller, muzzles larger, the anterior-most portion of the nasal bones generally flare upwards slightly, and the downward slope of the dorsal edge of the skull in front of the orbit is not as pronounced, producing a straighter profile. Compared with the most well known machairodont ''Smilodon'', commonly referred to as the "saber-toothed cat", the canines are much shorter, the facial portion again is much longer, and the teeth not reduced so far in number. Several machairodonts, namely ''Megantereon'', bear flanges on the mandible, which are very reduced in ''A. giganteus'' though characteristics of the mandible associated with the flanges are present, particularly the lateral flattening of the anterior portion of the mandible, creating a cross section more square than semi-circular. The dental formula for this specimen is .

Paleoecology

''Amphimachairodus'' was an inhabitant of woodlands and open floodplains as based on finds in Pikermi in Greece and Shanxi Province in China, indicating it had habitat preferences similar to modern lions in many respects. Specimens recovered from Turolian deposits indicate that the fauna living there was much the same, differing only by species in many cases.
Among the creatures it shared its environment with were bovids such as '' Parabos'', Lutung monkeys, the mastodon '' Anancus'', the rhino ''Aceratherium'', antelopes such as '' Tragoportax'' and ''Miotragocerus'' as well as gazelles and deer, a very large species of hyrax, early goats, various giraffes, camels such as ''Paracamelus'', the horse ''Hipparion'', a species of aardvark, the chalicothere '' Ancylotherium'' and the beaver-like '' Dipoides''.
Other carnivores it shared its territory with include the percrocutid '' Dinocrocuta'', the bear ''Agriotherium'', fellow machairodonts ''Metailurus'' and ''Paramachairodus'' and hyenas like ''Thalassictis''.

The larger herbivores were likely common prey for ''Amphimachairodus'', and it likely would have competed with ''Agriotherium'' for food, possibly yielding kills to the bear and possibly also stealing kills from hyenas such as ''Thalassictis'' and from ''Metailurus'' when the opportunity arose.

In North America, in places such as Coffee Ranch in Texas, ''Amphimachairodus coloradensis'' shared territory with ''Agriotherium'' as it had in Africa and Eurasia, but also shared territory with the feliform ''Barbourofelis'' and the canids ''Epicyon'' and '' Borophagus'', and herbivores like the camel ''Aepycamelus'', the pronghorn antelope ''Cosoryx'', horses like ''Neohipparion'' and ''Nannippus'', the peccary '' Prosthennops'' and rhinoceroses like ''Teleoceras''.

In the Djurab desert in northern Chad, ''Amphimachairodus kabir'' co-existed with fellow machairodonts '' Lokotunjailurus'', '' Tchadailurus'' and early representatives of the genus ''Megantereon''. In addition, animals such as crocodiles, three-toed horses, fish, monkeys, hippos, aardvarks, turtles, rodents, giraffes, snakes, antelopes, pigs, mongooses, foxes, hyenas, otters, honey badgers and the hominid ''Sahelanthropus'' dwelled here, providing ample food. Based on these and other fossils, it is theorized that the Djurab was once the shore of a lake, generally forested close to the shore with savannah-like areas some distance away. The great number of cat species in the environment indicates that there was significant prey and available niches for multiple species of large felids to coexist.Scientific American
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See also

* Big cat
* List of largest carnivorans
* List of largest prehistoric carnivorans
* Megafauna
* Miocene
* Quaternary extinction event

References

{{Taxonbar, from=Q16974134

Homotherini
Miocene mammals of Europe
Miocene mammals of Africa
Miocene mammals of Asia
Miocene mammals of North America
Fossil taxa described in 1929
Prehistoric carnivoran genera