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''Acis ionica'' is a
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
in the family
Amaryllidaceae The Amaryllidaceae are a family of herbaceous, mainly perennial and bulbous (rarely rhizomatous) flowering plants in the monocot order Asparagales. The family takes its name from the genus '' Amaryllis'' and is commonly known as the amaryl ...
, native from south-western Albania to western Greece. It was initially confused with what is now '' Acis valentina'', a species found in Spain near
Valencia Valencia ( va, València) is the capital of the autonomous community of Valencia and the third-most populated municipality in Spain, with 791,413 inhabitants. It is also the capital of the province of the same name. The wider urban area al ...
.


Description

''Acis ionica'' is a
bulb In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs du ...
ous plant with narrow leaves, 12–22 cm long but only 2–3 mm wide. The leaves appear after flowering, which is in the autumn. The flowering stalk ( scape) is usually 8–20 cm tall, with an
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed ...
composed of two to four flowers, occasionally only one or as many as six. The inflorescence is subtended by two spathes, usually extending to below the level of the highest flower. Each flower opens widely and is borne on a stem (
pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...
) about 16–28 mm long. The six
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s are white, generally 9–13 mm long. The outer three tepals have pointed tips. The pedicel lengthens after flowering. The black seeds are about 3 mm long.


Taxonomy

''Acis'' species were at one time placed in the genus ''Leucojum''. What is now considered to be ''Acis ionica'' was initially confused with ''Leucojum valentinum'' ('' Acis valentina''), a species found in Spain near
Valencia Valencia ( va, València) is the capital of the autonomous community of Valencia and the third-most populated municipality in Spain, with 791,413 inhabitants. It is also the capital of the province of the same name. The wider urban area al ...
. Doubts over this identification began around 1990, and both morphological and molecular evidence began to suggest that it was a distinct species. It was first described as a separate species in 2004 as ''Leucojum ionicum''. Separately, and on the basis of a different
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * Ty ...
, ''Acis ionica'' was described in 2006 by Pepy Bareka, Georgia Kamari and Dimitrios Phitos. In 2019,
Arne Strid P. Arne K. Strid (born March 7, 1943 in Kristianstad, Sweden) is a Swedish botanist and expert on Greek flora. Biography He studied botany, chemistry and genetics in the University of Lund and graduated in 1970. His doctorate was about an experi ...
treated both taxa as the same species based on the very limited differences between their characters. In order to transfer the first published name, ''Leucojum ionicum'', to the genus ''Acis'', Strid considered that it was necessary to use a replacement name (''
nomen novum In biological nomenclature, a ''nomen novum'' (Latin for "new name"), new replacement name (or replacement name, new substitute name, substitute name) is a scientific name that is created specifically to replace another scientific name, but only w ...
'') to avoid creating a
homonym In linguistics, homonyms are words which are homographs (words that share the same spelling, regardless of pronunciation), or homophones (equivocal words, that share the same pronunciation, regardless of spelling), or both. Using this definition, ...
of the already published ''Acis ionica''. For this purpose, Strid published the name ''Acis orientalis''. A different view is adopted by the ''
World Checklist of Selected Plant Families The World Checklist of Selected Plant Families (usually abbreviated to WCSP) is an "international collaborative programme that provides the latest peer reviewed and published opinions on the accepted scientific names and synonyms of selected pl ...
'', namely that the next published name, ''Acis ionica'', should be used, making Strid's replacement name superfluous and hence
illegitimate Legitimacy, in traditional Western common law, is the status of a child born to parents who are legally married to each other, and of a child conceived before the parents obtain a legal divorce. Conversely, ''illegitimacy'', also known as ...
. Although similar in appearance to ''A. valentina'', genetically it is as distinct as any other species of ''Acis''. Compared to ''A. valentina'', it has a thinner scape (flower stalk), broader inner
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s, and its flowers open more widely.


Distribution and habitat

''Acis ionica'' is found in western Greece, on the islands of
Zakynthos Zakynthos (also spelled Zakinthos; el, Ζάκυνθος, Zákynthos ; it, Zacinto ) or Zante (, , ; el, Τζάντε, Tzánte ; from the Venetian form) is a Greek island in the Ionian Sea. It is the third largest of the Ionian Islands. Za ...
,
Kefalonia Kefalonia or Cephalonia ( el, Κεφαλονιά), formerly also known as Kefallinia or Kephallenia (), is the largest of the Ionian Islands in western Greece and the 6th largest island in Greece after Crete, Euboea, Lesbos, Rhodes and Chio ...
and Lefkas, with populations on the nearby mainland of Greece and further north in coastal
Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic The Adriatic Sea () is a body of water separating the ...
(close to the city of
Vlorë Vlorë ( , ; sq-definite, Vlora) is the third most populous city of the Republic of Albania and seat of Vlorë County and Vlorë Municipality. Located in southwestern Albania, Vlorë sprawls on the Bay of Vlorë and is surrounded by the foot ...
). It is found on calcareous soils, in open, stony or rocky ground and on hillsides, usually not far from the sea and at altitudes below 350 m.


Notes


References

{{Taxonbar, from1=Q15517214, from2=Q21276517, from3=Q86428961 ionica Flora of Albania Flora of Greece Plants described in 2004