Achlys (plant)
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''Achlys'' is a small genus of flowering plants in the barberry family (
Berberidaceae The Berberidaceae are a family (biology), family of 18 genera of flowering plants commonly called the barberry family. This family is in the order (biology), order Ranunculales. The family contains about 700 known species, of which the majority ...
), which it shares with genera such as ''
Berberis ''Berberis'' (), commonly known as barberry, is a large genus of deciduous and evergreen shrubs from tall, found throughout temperate and subtropical regions of the world (apart from Australia). Species diversity is greatest in South America a ...
'' and ''
Vancouveria ''Vancouveria'' () is a small group of plants belonging to the Berberidaceae, barberry family described as a genus in 1834. The three plants in this genus are known generally as inside-out flowers, and they are endemic to the West Coast of the Un ...
''.Gilkey, Helen M. & Dennis, La Rea J. (2001). ''Handbook of Northwestern Plants''. Oregon State University Press. p. 161 It is named after the Greek legendary figure associated with shade and mist,
Achlys Achlys ( "mist"), in the Hesiodic '' Shield of Heracles'', is one of the figures depicted on Heracles' shield, perhaps representing the personification of sorrow. In Homer, ''achlys'' is the mist which fogs or blinds mortal eyes (often in dea ...
, because the plants grow in the shade.


Species

There are either two or three species, depending on the authority. ''Achlys triphylla'' and ''Achlys californica'' are both native to western North America. Another ''Achlys'' is found in Japan: some authorities treat this as a subspecies of ''A. triphylla'', while others, especially in older treatments, call this ''Achlys japonica''. Still others consider ''A. triphylla'' and ''A. californica'' too similar to be separate species. The common names for these plants include vanilla leaf (sometimes written as vanilla-leaf or vanillaleaf, depending on the taxonomist or flora), deer's foot and sweet after death, referring to the vanilla scent of its crushed leaves.Pojar & MacKinnon (2004). ''Plants of the Pacific Northwest Coast'' (Revised Edition). Lone Pine Publishing. p. 312
The Plant List The Plant List was a list of botanical names of species of plants created by the Royal Botanic Gardens, Kew and the Missouri Botanical Garden and launched in 2010. It was intended to be a comprehensive record of all known names of plant specie ...
recognizes two species, with ''A. californica'' regarded as a subspecies of ''A. triphylla''


Description

''Achlys triphylla'', known in western North America as vanillaleaf, is an erect perennial plant that sprouts from a creeping
rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...
. Leaves are long-petioled and palmately divided into three leaflets. Flowers are small and lack sepals and petals, but instead have long showy white
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s that form single erect spikes. The leaflets give a great hint to the identification of the plant: bend back the middle leaflet and you have an upside-down set of moose antlers. Alternatively, bend back the two side leaflets and you have a goose or deer foot (hence the common name). In the Pacific Northwest, ''Achlys triphylla'' is ubiquitous in moist shady forests west of the Cascades at low to middle elevations from Vancouver island and southern
British Columbia British Columbia is the westernmost Provinces and territories of Canada, province of Canada. Situated in the Pacific Northwest between the Pacific Ocean and the Rocky Mountains, the province has a diverse geography, with rugged landscapes that ...
south to northern
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
. The plants are spaced widely on the rhizomes, but often overlap in large networks that result in carpets of ''Achlys'' that dominate the near-surface understory. ''Achlys'' seems to prefer moist soil, so at middle to higher elevations it is easier to find them along streambanks or well-shaded ravines. File:Achlys japonica 5.JPG, ''Achlys japonica'' flowers File:Achlys triphylla 0973.JPG, Covered with fine hairs, ''Achlys triphylla'' achenes are concave toward stem and fleshy. File:Achlys triphylla 15379.JPG, ''Achlys triphylla'' at Standup Creek Trail 1369 Wenatchee National Forest File:Achlys japonica 6.JPG, ''Achlys japonica'' growing in Kitakami Mountains, Iwate prefecture, Japan


Insect repellent

When dried properly, the plants are strongly aromatic and smell of vanilla. Besides serving as an excellent tent
air freshener Air fresheners are products designed to reduce unwanted odors in indoor spaces, to introduce pleasant fragrances, or both. They typically emit fragrance to mask odors but may use other methods of action such as absorbing, bonding to, or chemically ...
, ''Achlys'' has been used by native tribes of southern British Columbia as an
insect repellent An insect repellent (also commonly called "bug spray" or "bug deterrent") is a substance applied to the skin, clothing, or other surfaces to discourage insects (and arthropods in general) from landing or climbing on that surface. Insect repellent ...
. The dried leaves were hung in bunches in doorways to ward off flies and
mosquitoe Mosquitoes, the Culicidae, are a family of small flies consisting of 3,600 species. The word ''mosquito'' (formed by '' mosca'' and diminutive ''-ito'') is Spanish and Portuguese for ''little fly''. Mosquitoes have a slender segmented body, ...
s, and it's not unheard of for naturalists to rub the dried or even fresh leaves on exposed skin when hiking the Olympics or Cascades during the summer mosquito season. Japanese ''Achlys'' are quite similar to those found in western North America.


References


External links

*
Jepson Manual eFlora (TJM2) treatment of ''Achlys''
{{Taxonbar, from=Q4673829 Berberidaceae Berberidaceae genera Flora of British Columbia Flora of California Flora of the West Coast of the United States Insect repellents Plant toxin insecticides