''Abronia umbellata'' (pink sand verbena) is a flowering

annual plant An annual plant is a plant that completes its life cycle, from germination to the production of seeds, within one growing season, and then dies. Globally, 6% of all plant species and 15% of herbaceous plants (excluding trees and shrubs) are ...

which is

native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...

to western North America. Other common names include beach sand verbena and purple sand verbena.

Distribution and habitat

This plant is generally found in sandy, well-drained

soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...

in areas with low precipitation, it can become a striking carpet-like

groundcover Groundcover or ground cover is any plant that grows low over an area of ground, which protects the topsoil from erosion and drought. In a terrestrial ecosystem, the ground cover forms the layer of vegetation below the shrub layer known as the ...

in undisturbed areas after winter rains. Pink sand verbena tolerates seaside conditions and is found on the west coast of North America from

British Columbia, Canada British Columbia is the westernmost province of Canada. Situated in the Pacific Northwest between the Pacific Ocean and the Rocky Mountains, the province has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beach ...

Baja California Baja California, officially the Free and Sovereign State of Baja California, is a state in Mexico. It is the northwesternmost of the 32 federal entities of Mexico. Before becoming a state in 1952, the area was known as the North Territory of B ...

Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...

. Sand verbena is typically found on beaches and sand dunes, below the

coastal sage scrub Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland plant community of the California coastal sage and chaparral subecoregion, found in coastal California and northwestern coastal Baja California. It is ...

, blooming throughout most of the year. It is listed as endangered by the State of Oregon.

Description

''Abronia umbellata'' is a prostrate annual with thick, succulent leaves (leaves occur few to many and are slender, ovate to diamond-shape with stems as long as leaf blades, stems are often hairy) and pink to purple colored flowers with white centers. Flowers occur in clusters subtended by 5-8 lanceolate bracts. The flowers do not have

petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...

s, but the calyx lobes are cleft giving the appearance of 10-16 petals. The limbs of the

perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower. It is a structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepal ...

is bright colored sometimes to purplish magenta and the tube can be green or red but always-glandular

pubescent The adjective pubescent may describe: * people or animals undergoing puberty * plants that are hairy, covered in trichomes * insects that are covered in setae In biology, setae (; seta ; ) are any of a number of different bristle- or hair-lik ...

. The tube includes one pistil and three stamens. ''A. umbellata'' frequently hybridizes with other species of ''Abronia'', including '' A. maritima''. Its flower is fragrant at night and attracts moths. The foliage can be deciduous based on environmental stress. This plant is sometimes used in California in native plant gardening.

History in Europe

Originally described in 1793 by the French botanist

Jean-Baptiste Lamarck Jean-Baptiste Pierre Antoine de Monet, chevalier de Lamarck (1 August 1744 – 18 December 1829), often known simply as Lamarck (; ), was a French naturalist, biologist, academic, and soldier. He was an early proponent of the idea that biologi ...

, ''Abronia umbellata'' was collected in 1786 from

Monterey, California Monterey ( ; ) is a city situated on the southern edge of Monterey Bay, on the Central Coast (California), Central Coast of California. Located in Monterey County, California, Monterey County, the city occupies a land area of and recorded a popu ...

by the gardener Jean Nicolas Collignon of the French La Pérouse expedition, which had stopped at the capital of

Alta California Alta California (, ), also known as Nueva California () among other names, was a province of New Spain formally established in 1804. Along with the Baja California peninsula, it had previously comprised the province of , but was made a separat ...

as part of a journey of scientific exploration spanning the Pacific Ocean. While Collignon and his shipmates perished in a wreck near

Vanikoro Vanikoro (sometimes wrongly named ''Vanikolo'') is an island in the Santa Cruz Islands, Santa Cruz group, located to the Southeast of the main Santa Cruz group. It is part of the Temotu Province of Solomon Islands. The name ''Vanikoro'' is alway ...

in the

Solomon Islands Solomon Islands, also known simply as the Solomons,John Prados, ''Islands of Destiny'', Dutton Caliber, 2012, p,20 and passim is an island country consisting of six major islands and over 1000 smaller islands in Melanesia, part of Oceania, t ...

, some of his collection had previously been shipped back to France during a stop at the Portuguese-held

Macao Macau or Macao is a special administrative region of the People's Republic of China (PRC). With a population of about people and a land area of , it is the most densely populated region in the world. Formerly a Portuguese colony, the ter ...

, including the seeds of ''A. umbellata''. They were planted at the

Jardin des Plantes The Jardin des Plantes (, ), also known as the Jardin des Plantes de Paris () when distinguished from other ''jardins des plantes'' in other cities, is the main botanical garden in France. Jardin des Plantes is the official name in the present da ...

in Paris, and Lamarck eventually named their descendants ''A. umbellata'', making this species the first Californian flower that does not occur outside of western North America that was described in the scientific fashion of

Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

References

External links

Jepson Manual treatment – ''Ambronia umbellata''United States Department of Agriculture Plants Profile: ''Abronia umbellata''''Abronia umbellata'' – Calphotos Photo Gallery, University of CaliforniaLady Bird Johnson Wildflower Center, University of Texas at Austin: ''Abronia umbellata''
{{Taxonbar, from=Q4055083 umbellata Flora of the West Coast of the United States Flora of British Columbia Flora of Baja California Endangered flora of the United States Plants described in 1793