
The 19th century in ichnology refers to advances made between the years
1800
As of March 1 ( O.S. February 18), when the Julian calendar acknowledged a leap day and the Gregorian calendar did not, the Julian calendar fell one day further behind, bringing the difference to 12 days until February 28 ( O.S. February 16), ...
and
1899
Events January 1899
* January 1
** Spanish rule ends in Cuba, concluding 400 years of the Spanish Empire in the Americas.
** Queens and Staten Island become administratively part of New York City.
* January 2 –
**Bolivia sets up a c ...
in the scientific study of
trace fossils, the preserved record of the
behavior
Behavior (American English) or behaviour (British English) is the range of actions and mannerisms made by individuals, organisms, systems or artificial entities in some environment. These systems can include other systems or organisms as wel ...
and
physiological processes of ancient life forms, especially
fossil footprints. The 19th century was notably the first century in which fossil footprints received scholarly attention. British
paleontologist William Buckland performed the first true scientific research on the subject during the early
1830s
The 1830s (pronounced "eighteen-thirties") was a decade of the Gregorian calendar that began on January 1, 1830, and ended on December 31, 1839.
In this decade, the world saw a rapid rise of imperialism and colonialism, particularly in Asia an ...
.
A slab of
Permian-aged
sandstone had been discovered in Scotland which preserved a series of unusual footprints. After acquiring the specimen, Buckland experimented with modern animals to ascertain the trackmaker and concluded that the Scottish footprints were made by
tortoises.
Later in the century famed advocate of evolution
Thomas Henry Huxley would refute this attribution and these footprints, called ''
Chelichnus'', would remain without an identified trackmaker until scientists recognized that they were actually made by an evolutionary
precursor to mammals.
The 1830s also saw the discovery and investigation of unusual
hand
A hand is a prehensile, multi-fingered appendage located at the end of the forearm or forelimb of primates such as humans, chimpanzees, monkeys, and lemurs. A few other vertebrates such as the koala (which has two opposable thumbs on each "h ...
-shaped footprints from
Triassic rocks in
Germany that were later named ''
Chirotherium''. The identification of the ''Chirotherium'' trackmaker proved elusive and suggestions from researchers included everything from
monkeys
Monkey is a common name that may refer to most mammals of the infraorder Simiiformes, also known as the simians. Traditionally, all animals in the group now known as simians are counted as monkeys except the apes, which constitutes an incomple ...
to giant
toads and
kangaroos.
''Chirotherium'' proved to be an enduring ichnological mystery that would not be solved until long into the 20th century.
Some of the most important ichnological research of the 19th century occurred across the
Atlantic in the United States.
Dinosaur footprints
A fossil track or ichnite (Greek "''ιχνιον''" (''ichnion'') – a track, trace or footstep) is a fossilized footprint. This is a type of trace fossil. A fossil trackway is a sequence of fossil tracks left by a single organism. Over the year ...
were first discovered there in
1802
Events
January–March
* January 5 – Thomas Bruce, 7th Earl of Elgin, British ambassador to the Ottoman Empire, begins removal of the Elgin Marbles from the Parthenon in Athens, claiming they were at risk of destruction during the Ot ...
when a
Massachusetts farm boy stumbled upon
bird-like footprints in sandstone that the local
clergy mistakenly attributed to the
raven
A raven is any of several larger-bodied bird species of the genus ''Corvus''. These species do not form a single taxonomic group within the genus. There is no consistent distinction between "crows" and "ravens", common names which are assigned t ...
that
Noah
Noah ''Nukh''; am, ኖህ, ''Noḥ''; ar, نُوح '; grc, Νῶε ''Nôe'' () is the tenth and last of the pre-Flood patriarchs in the traditions of Abrahamic religions. His story appears in the Hebrew Bible (Book of Genesis, chapters 5– ...
released from his
ark during the
Biblical Flood.
The region's footprints came to the attention of scholars during the mid 1830s when further bird-like dinosaur tracks were discovered elsewhere
in the state. These became the lifelong preoccupation of prominent ichnologist
Edward Hitchcock
Edward Hitchcock (May 24, 1793 – February 27, 1864) was an American geologist and the third President of Amherst College (1845–1854).
Life
Born to poor parents, he attended newly founded Deerfield Academy, where he was later principal, ...
. Hitchcock thought the tracks were made by giant
flightless birds.
Late in the 19th century prisoners
in Nevada discovered a major
Ice Age track site at what was once an ancient
lake shore. Many of the trackmakers were familiar animals like
mammoths or even more modern animals like
deer and
wolves, but this track site also seemed to preserve the tracks of a
sandal-wearing
giant.
The tracks received significant scholarly and popular attention like satire by
Mark Twain
Samuel Langhorne Clemens (November 30, 1835 – April 21, 1910), known by his pen name Mark Twain, was an American writer, humorist, entrepreneur, publisher, and lecturer. He was praised as the "greatest humorist the United States has p ...
who attributed the giant tracks to primitive Nevadan legislators. However, the true identity of the "giant" trackmaker was recognized by paleontologists
Joseph Le Conte and
Othniel Charles Marsh as a giant
ground sloth
Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. The term is used to refer to all extinct sloths because of the large size of the earliest forms discovered, compared to existing tree sloths. The Caribbe ...
, possibly of the genus ''
Mylodon''.
19th century
1800s
1802
Events
January–March
* January 5 – Thomas Bruce, 7th Earl of Elgin, British ambassador to the Ottoman Empire, begins removal of the Elgin Marbles from the Parthenon in Athens, claiming they were at risk of destruction during the Ot ...
* ''Spring:'' A boy named Pliny Moody uncovered a piece of
sandstone with mysterious three-toed tracks about 30 cm (1 foot) long while plowing in his father's fields near
South Hadley, Massachusetts
South Hadley (, ) is a town in Hampshire County, Massachusetts, United States. The population was 18,150 at the 2020 census. It is part of the Springfield, Massachusetts Metropolitan Statistical Area.
South Hadley is home to Mount Holyoke Colleg ...
. The local clergy thought the tracks had been left by the
raven
A raven is any of several larger-bodied bird species of the genus ''Corvus''. These species do not form a single taxonomic group within the genus. There is no consistent distinction between "crows" and "ravens", common names which are assigned t ...
that
Noah
Noah ''Nukh''; am, ኖህ, ''Noḥ''; ar, نُوح '; grc, Νῶε ''Nôe'' () is the tenth and last of the pre-Flood patriarchs in the traditions of Abrahamic religions. His story appears in the Hebrew Bible (Book of Genesis, chapters 5– ...
sent out from the
ark to look for dry land during the
Biblical Flood.
1820s
1820s
File:1820s collage.jpg, 420x420px, From top left, clockwise: Ludwig van Beethoven re-emerged as a popular composer during this decade, when his iconic Symphony No. 9 is first performed in Vienna in 1824. The First Industrial Revolution achieves ...
* A slab of
Permian sandstone preserving 24 small footprints came into the possession of the
Scottish
Scottish usually refers to something of, from, or related to Scotland, including:
*Scottish Gaelic, a Celtic Goidelic language of the Indo-European language family native to Scotland
*Scottish English
*Scottish national identity, the Scottish ide ...
Reverend
Henry Duncan. Duncan visited the quarry where his slab was originally excavated in
Corncockle Muir
''Agrostemma'' is a genus of annual plants in the family Caryophyllaceae, containing the species known as corncockles. Its best-known member is ''A. githago'', the common corncockle, which is a native of Europe. The species is a weed of cereals ...
to see if he could find more of the fascinating impressions and successfully recovered more of them. He notified leading paleontologist
William Buckland of
Oxford University about his discovery.
Late
1820s
File:1820s collage.jpg, 420x420px, From top left, clockwise: Ludwig van Beethoven re-emerged as a popular composer during this decade, when his iconic Symphony No. 9 is first performed in Vienna in 1824. The First Industrial Revolution achieves ...
* Fossil footprints were reported from
Cheshire
Cheshire ( ) is a ceremonial and historic county in North West England, bordered by Wales to the west, Merseyside and Greater Manchester to the north, Derbyshire to the east, and Staffordshire and Shropshire to the south. Cheshire's county t ...
, England in
Triassic rocks known at the time as the
New Red Sandstone. This is the earliest written record of fossil footprints now referred to the
ichnogenus ''
Chirotherium''.
1828
Events
January–March
* January 4 – Jean Baptiste Gay, vicomte de Martignac succeeds the Comte de Villèle, as Prime Minister of France.
* January 8 – The Democratic Party of the United States is organized.
* January 22 – Arthu ...
* A newspaper article was published that discussed fossil footprints, and is now regarded as the earliest written record of the subject.
1830s
1831
Events
January–March
* January 1 – William Lloyd Garrison begins publishing '' The Liberator'', an anti-slavery newspaper, in Boston, Massachusetts.
* January 10 – Japanese department store, Takashimaya in Kyoto establ ...
* Buckland published the first scientific description of fossil footprints about the tracks discovered at Corncockle Muir. He attributed the footprints to ancient
tortoises because after having various modern reptiles walk over stretches of
pie
A pie is a baked dish which is usually made of a pastry dough casing that contains a filling of various sweet or savoury ingredients. Sweet pies may be filled with fruit (as in an apple pie), nuts ( pecan pie), brown sugar ( sugar pie), swe ...
crust
dough, the tracks left by tortoises most closely resembled those from the Permian sandstone.
1834
Events
January–March
* January – The Wilmington and Raleigh Railroad is chartered in Wilmington, North Carolina.
* January 1 – Zollverein (Germany): Customs charges are abolished at borders within its member states.
* January 3 � ...

* A man named
Helmut Barth was building a garden house in
Hildburghausen, Germany when he discovered strange, hand-shaped tracks in the sandstone he was using in the construction.
Barth's discovery would be named ''Chirotherium'' by
Johan Jacob Kaup.
1835
Events
January–March
* January 7 – anchors off the Chonos Archipelago on her second voyage, with Charles Darwin on board as naturalist.
* January 8 – The United States public debt contracts to zero, for the only time in history.
...
* While the streets of
Greenfield, Massachusetts were being paved, locals noticed footprints impressed in the stone. The townspeople thought the tracks were left by
turkeys.
They informed James Deane, a local doctor and naturalist about the footprints. Deane found the tracks intriguing and wrote to another local scholar,
Edward B. Hitchcock about the find.
Hitchcock spent the rest of the summer investigating the local footprints fossils.
1836
Events
January–March
* January 1 – Queen Maria II of Portugal marries Prince Ferdinand Augustus Francis Anthony of Saxe-Coburg-Gotha.
* January 5 – Davy Crockett arrives in Texas.
* January 12
** , with Charles Darwin on board, r ...
* Edward Hitchcock published the results of his research into the fossil footprints. He thought the trackmakers were large
flightless birds.
1840s
1842
Events
January–March
* January
** Michael Alexander takes office, as the first appointee to the Anglican-German Bishopric in Jerusalem.
** American medical student William E. Clarke of Berkshire Medical College becomes the first pe ...
*
Sir Richard Owen supported Buckland's interpretation of the Corncockle Muir tracks as tortoise tracks and named them ''
Testudo duncani''.
1843
Events January–March
* January
** Serial publication of Charles Dickens's novel ''Martin Chuzzlewit'' begins in London; in the July chapters, he lands his hero in the United States.
** Edgar Allan Poe's short story "The Tell-Tale Heart" ...
* James Deane published the results of his own investigations into the fossil footprints of Massachusetts.
1845
Events
January–March
* January 10 – Elizabeth Barrett receives a love letter from the younger poet Robert Browning; on May 20, they meet for the first time in London. She begins writing her ''Sonnets from the Portuguese''.
* January 23 ...
* Hitchcock named the ichnogenus ''
Eubrontes''.
1846
Events
January–March
* January 5 – The United States House of Representatives votes to stop sharing the Oregon Country with the United Kingdom of Great Britain and Ireland, United Kingdom.
* January 13 – The Milan–Venice railway' ...
*
Tagart reported dinosaur tracks in England that would later come to be attributed to ''
Iguanodon''.
1847
Events
January–March
* January 4 – Samuel Colt sells his first revolver pistol to the U.S. government.
* January 13 – The Treaty of Cahuenga ends fighting in the Mexican–American War in California.
* January 16 – John C. Frémont ...
* A man surnamed Cotta wrote a letter including the first documented mention of the many Permian tracks preserved in the "
Rotliegendes" of central Germany's
Thuringian forest.
"Rotliegendes" is German for "red layers" referring to a Permian sandstone layer rich in rusted
iron minerals known elsewhere as the "New Red Sandstone".
The tracks Cotta reported were later named ''
Saurichnites cottae'' in his honor.
1848
1848 is historically famous for the wave of revolutions, a series of widespread struggles for more liberal governments, which broke out from Brazil to Hungary; although most failed in their immediate aims, they significantly altered the polit ...
* Hitchcock name the ichnogenus ''
Anomoepus''.
1850s
1850
Events
January–June
* April
** Pope Pius IX returns from exile to Rome.
** Stephen Foster's parlor ballad "Ah! May the Red Rose Live Alway" is published in the United States.
* April 4 – Los Angeles is incorporated as a cit ...
*
Sir William Jardine
Sir William Jardine, 7th Baronet of Applegarth FRS FRSE FLS FSA (23 February 1800 – 21 November 1874) was a Scottish naturalist. He is known for his editing of a long series of natural history books, ''The Naturalist's Library''.
Life a ...
argued against Owen's referral of the Corncockle Muir "tortoise" footprints to ''
Testudo'' because the name applied to a specific group of modern
turtles rather than to footprints. He coined the name ''
Chelichnus'', meaning "turtle track" to replace Owen's use of ''Testudo'', but preserved the specific epithet "''duncani''".
1851
Events
January–March
* January 11 – Hong Xiuquan officially begins the Taiping Rebellion.
* January 15 – Christian Female College, modern-day Columbia College, receives its charter from the Missouri General Assembly.
...
* More Permian ''Chelichnus'' tracks were discovered in the highlands of Scotland, not far from
Cummingstone.
*
Samuel H. Beckles began publishing research on the dinosaur footprints from the
Wealden, although he did not recognize their dinosaurian origins.
1852
Events
January–March
* January 14 – President Louis-Napoléon Bonaparte proclaims a new constitution for the French Second Republic.
* January 15 – Nine men representing various Jewish charitable organizations come tog ...
* Beckles continued to publish research on the dinosaur footprints from the Wealden.
1854
Events
January–March
* January 4 – The McDonald Islands are discovered by Captain William McDonald aboard the ''Samarang''.
* January 6 – The fictional detective Sherlock Holmes is perhaps born.
* January 9 – The Teut ...
* Beckles continued to publish research on the dinosaur footprints from the Wealden, referring to them as Ornithoidichnites following the nomenclature devised by Edward Hitchcock for some American tracks. Despite his use of a term implying an avian trackmaker, Beckles admitted that he did not know what kind of animals made the tracks.
1858
Events
January–March
* January –
**Benito Juárez (1806–1872) becomes Liberal President of Mexico. At the same time, conservatives install Félix María Zuloaga (1813–1898) as president.
**William I of Prussia becomes regent f ...
* Edward Hitchcock published a summary of his research into the fossil footprints of the
Connecticut Valley area. He continued to attribute the tracks to large flightless birds that he named their footprints "ornithichnites", meaning "stone bird footprints". He divided the trackmakers into two groups, the leptodactylous birds with narrow toes and the pachydactylous birds with thick toes. He also described seven new ichnospecies for the tracks he studied.
He also described the ichnogenus ''
Grallator''.
1860s
1860
Events
January–March
* January 2 – The discovery of a hypothetical planet Vulcan is announced at a meeting of the French Academy of Sciences in Paris, France.
* January 10 – The Pemberton Mill in Lawrence, Massachusett ...
* A British scientists surnamed Williamson interpreted the ''Chirotherium'' trackmaker as a
crocodile
Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to inclu ...
.
1862
Events
January–March
* January 1 – The United Kingdom annexes Lagos Island, in modern-day Nigeria.
* January 6 – French intervention in Mexico: French, Spanish and British forces arrive in Veracruz, Mexico.
* January ...
*
Oppel Oppel is a surname. Notable people with the surname include:
* Albert Oppel (1831–1865), German paleontologist
* Dorsum Oppel, a wrinkle ridge in Mare Crisium on the Moon
* Kenneth Oppel (born 1967), Canadian author
*Nicolaus Michael Oppel (1782� ...
interpreted some tracks from the
Solnhofen lithographic limestone as the tracks of
pterosaurs
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
.
1864
Events
January–March
* January 13 – American songwriter Stephen Foster ("Oh! Susanna", "Old Folks at Home") dies aged 37 in New York City, leaving a scrap of paper reading "Dear friends and gentle hearts". His parlor song " ...
* Edward Hitchcock died.
1865
Events
January–March
* January 4 – The New York Stock Exchange opens its first permanent headquarters at Broad Street (Manhattan), 10-12 Broad near Wall Street, in New York City.
* January 13 – American Civil War : Sec ...
* A posthumous "supplement" to Hitchcock's monograph on the Connecticut Valley tracks was published.
* 1860: Some English dinosaur footprints were recognized as ''Iguanodon'' tracks. They were the first dinosaur tracks to be recognized as belonging to an individual genus.
1870s
1877
Events
January–March
* January 1 – Queen Victoria is proclaimed ''Empress of India'' by the ''Royal Titles Act 1876'', introduced by Benjamin Disraeli, the Prime Minister of the United Kingdom .
* January 8 – Great Sio ...
*
Thomas Henry Huxley argued against Buckland and Owen's attribution of ''Chelichnus duncani'' to ancient tortoises, instead concluding that it was impossible to identify the trackmaker with the knowledge of time.
1879
Events January–March
* January 1 – The Specie Resumption Act takes effect. The United States Note is valued the same as gold, for the first time since the American Civil War.
* January 11 – The Anglo-Zulu War begins.
* Janu ...
* Some Welsh dinosaur tracks that had been previously displayed in front of the Jolly Sailor inn in
Newton
Newton most commonly refers to:
* Isaac Newton (1642–1726/1727), English scientist
* Newton (unit), SI unit of force named after Isaac Newton
Newton may also refer to:
Arts and entertainment
* ''Newton'' (film), a 2017 Indian film
* Newton ( ...
were acquired by the
Cardiff Museum.
*
T. H. Thomas
Thomas Henry Thomas (31 March 1839 – 9 July 1915) was a Welsh artist particularly active in Cardiff. He was also interested in botany, geology, history, and archaeology which were often the subjects of his art works. He was a Fellow of the Roya ...
reported the Welsh dinosaur footprints to the scientific literature and noted their similarity to the "Ornithichnites" of Connecticut.
*
W. J. Sollas independently published a report of the Welsh dinosaur footprints.
1880s
Early 1880s
* Inmates of
Nevada State Prison uncovered a large
Pleistocene fossil track site while excavating sandstone. The track sites was a
lakeshore 50,000 years ago where familiar
Ice Age animals like birds,
deer,
mammoths, and
wolves left behind their footprints. However, ten of the roughly 50 trails seemed to have been left by an even stranger trackmaker; a sandaled
giant.
1880
Events
January–March
* January 22 – Toowong State School is founded in Queensland, Australia.
* January – The international White slave trade affair scandal in Brussels is exposed and attracts international infamy.
* February � ...
* George Le Mesle and Pierre Peron discovered dinosaur footprints in Algeria. These were the first fossil dinosaur tracks to be discovered in Africa.
1884
Events
January–March
* January 4 – The Fabian Society is founded in London.
* January 5 – Gilbert and Sullivan's ''Princess Ida'' premières at the Savoy Theatre, London.
* January 18 – Dr. William Price atte ...
* Large
theropod footprints were reported in
Late Jurassic rocks at
Cabo Mondego,
Portugal. These may have been the first European Late Jurassic dinosaur footprints to be documented in the scientific literature.~152~
*
W. P. Blake reported the fossil footprints discovered at the prison in
Carson City, Nevada to the scientific literature.
*
Mark Twain
Samuel Langhorne Clemens (November 30, 1835 – April 21, 1910), known by his pen name Mark Twain, was an American writer, humorist, entrepreneur, publisher, and lecturer. He was praised as the "greatest humorist the United States has p ...
wrote the satirical "The Carson Fossil Footprints" attributing the purported giant tracks discovered there to primitive members of the
territorial legislature.
1886
Events
January–March
* January 1 – Upper Burma is formally annexed to British Burma, following its conquest in the Third Anglo-Burmese War of November 1885.
* January 5– 9 – Robert Louis Stevenson's novella ''Strange ...
* A Mr. C. Pooley discovered a small five toed fossil footprint in
Oxfordshire
Oxfordshire is a ceremonial and non-metropolitan county in the north west of South East England. It is a mainly rural county, with its largest settlement being the city of Oxford. The county is a centre of research and development, primarily ...
, England, which preserved in a
Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
stratigraphic unit called the
Stonesfield Slate
Stonesfield is a village and civil parish about north of Witney in Oxfordshire, and about 10 miles (17 km) north-west of Oxford. The village is on the crest of an escarpment. The parish extends mostly north and north-east of the village, ...
.
*
John Eyerman discovered two slabs of rock preserving fossil dinosaur footprints near
Milford Milford may refer to:
Place names Canada
* Milford (Annapolis), Nova Scotia
* Milford (Halifax), Nova Scotia
* Milford, Ontario
England
* Milford, Derbyshire
* Milford, Devon, a place in Devon
* Milford on Sea, Hampshire
* Milford, Shro ...
,
New Jersey. The trackmaker was probably a small, quadrupedal
ornithischian dinosaur.
1889
Events
January–March
* January 1
** The total solar eclipse of January 1, 1889 is seen over parts of California and Nevada.
** Paiute spiritual leader Wovoka experiences a vision, leading to the start of the Ghost Dance movement in the ...
*
Addison Coffin drew up a map of a large portion of the Pleistocene Carson City tracks.
* Dinosaur footprints were discovered near
Goldsboro, Pennsylvania
Goldsboro (formerly Goldsborough) is a borough in York County, Pennsylvania, United States. The population was 930 at the 2020 census.
For historical reasons, the post office in Goldsboro is named "Etters", although there is no incorporated pl ...
. These tracks are now classified in the ichnogenus ''
Atreipus''.
1890s
1890s
The 1890s (pronounced "eighteen-nineties") was a decade of the Gregorian calendar that began on January 1, 1890, and ended on December 31, 1899.
In the United States, the 1890s were marked by a severe economic depression sparked by the Panic of ...
* Middle Jurassic dinosaur tracks were discovered on the
Yorkshire coast of England.
1894
Events January–March
* January 4 – A military alliance is established between the French Third Republic and the Russian Empire.
* January 7 – William Kennedy Dickson receives a patent for motion picture film in the United S ...
*
Othniel Charles Marsh described the new ichnogenus ''
Limnopus
''Limnopus'' is an ichnogenus of ancient tetrapod footprint. Its footprints have been found in Moscovian aged-rocks situated in Alverley, Shropshire, England, Colorado and West Virginia. The ''Limnopus'' tracks were probably made by ''Diadectes ...
'' for
Carboniferous
The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
tracks from the
coal beds of
Kansas.
1895
Events
January–March
* January 5 – Dreyfus affair: French officer Alfred Dreyfus is stripped of his army rank, and sentenced to life imprisonment on Devil's Island.
* January 12 – The National Trust for Places of Histor ...
* A geology professor named
James A. Mitchell discovered some small ''Grallator'' tracks in the Late Triassic
Gettysburg Formation of
Maryland. These are the first and only known dinosaur tracks in the state.
See also
*
History of paleontology
The history of paleontology traces the history of the effort to understand the history of life on Earth by studying the fossil record left behind by living organisms. Since it is concerned with understanding living organisms of the past, paleonto ...
**
Timeline of paleontology
Timeline of paleontology
Antiquity – 16th century
* 6th century B.C. — The pre-Socratic Greek philosopher Xenophanes of Colophon argues that fossils of marine organisms show that dry land was once under water.
* 4th century B.C. � ...
*
Timeline of ichnology
**
20th century in ichnology
Footnotes
References
*
*
*
*
External links
* * {{Wikisource-inline, Ichnology, Ichnology
19th century in paleontology
Ichnology
Ichnology