''Rubus hawaiensis'', also called the ''Ākala'', is one of two species (with '' R. macraei'') commonly known as Hawaiian raspberry, endemic to

Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...

. It is found on the islands of

Kauai Kauai (), anglicized as Kauai ( or ), is one of the main Hawaiian Islands. It has an area of 562.3 square miles (1,456.4 km2), making it the fourth-largest of the islands and the 21st-largest island in the United States. Kauai lies 73 m ...

Molokai Molokai or Molokai ( or ; Molokaʻi dialect: Morotaʻi ) is the fifth most populated of the eight major islands that make up the Hawaiian Islands archipelago in the middle of the Pacific Ocean. It is 38 by 10 miles (61 by 16 km) at its g ...

Maui Maui (; Hawaiian language, Hawaiian: ) is the second largest island in the Hawaiian archipelago, at 727.2 square miles (1,883 km2). It is the List of islands of the United States by area, 17th-largest in the United States. Maui is one of ...

O'ahu Oahu (, , sometimes written Oahu) is the third-largest and most populated island of the Hawaiian Islands and of the U.S. state of Hawaii. The state capital, Honolulu, is on Oahu's southeast coast. The island of Oahu and the uninhabited Northwe ...

, and

in mesic to wet forest at elevations of . In most areas it is not common, but in some places (such as the upper Koolau Gap in

Haleakalā Haleakalā (; Hawaiian: ), or the East Maui Volcano, is a massive, active shield volcano that forms more than 75% of the Hawaiian Islands, Hawaiian Island of Maui. The western 25% of the island is formed by another volcano, Mauna Kahalawai, als ...

and Laupāhoehoe Natural Area Reserve) it can be a dominant member of the understory vegetation. Although similar to the other Hawaiian species, ''

Rubus macraei ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. I ...

'', the two are believed to be derived from separate dispersals to Hawaii.

Description

''Rubus hawaiensis'' is a

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

, typically growing as a clump of erect or (when longer) arching canes, long. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are compound, with three leaflets. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

is red, large (up to long and wide), and edible but not often eaten, as it is sour and somewhat bitter. Although frequently described as prickle-free ("thornless"), and often used as an example of loss of defenses in island plants, most plants do have thin prickles at least when small. As the cane grows the outer layer of bark usually sheds, taking the prickles with it. Interest in breeding "thornless" varieties of edible raspberries (possibly even with distantly related species since most ''Rubus'' readily hybridize) has led to the introduction of several species of continental ''Rubus'' species which have since escape cultivation and become pests. These include the yellow Himalayan raspberry, ''

Rubus ellipticus ''Rubus ellipticus'', commonly known as ainselu, golden evergreen raspberry, golden Himalayan raspberry, or yellow Himalayan raspberry, is an Asian species of thorny fruiting shrub in the rose family. Its native range stretches from the Indian su ...

'', and the Florida prickly blackberry, '' R. argutus.''

Impact

The presence of invasive alien ''Rubus'' species along with two native species has led to a debate on

biological control Biological control or biocontrol is a method of controlling pests, whether pest animals such as insects and mites, weeds, or pathogens affecting animals or plants by using other organisms. It relies on predation, parasitism, herbivory, or o ...

. Specifically, whether an agent that might be able to control the alien species should be released even if it may have serious impacts on the native species, if the latter are not part of a major evolutionary diversification and not a major part of most ecosystems.

Mythology

This berry is believed to be the land counterpart to the '' limu kala'' both appearing in the first period of creation (''wā'') as mentioned in the

Kumulipo In Hawaiian religion, the Kumulipo is the creation chant, first recorded in the 18th century. It also includes a genealogy of the members of Hawaiian royalty and was created in honor of Kalaninuiamamao and passed down orally to his daughter Alapa ...

References

External links

Hawaiian Native Plant Genera - Rubus
Photographs of ''R. hawaiensis''. {{Taxonbar, from=Q3017550 hawaiensis Endemic flora of Hawaii Biota of Hawaii (island) Biota of Lanai Biota of Maui Biota of Molokai ~ Plants described in 1858 Flora without expected TNC conservation status