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Tomistomines
Tomistominae is a subfamily of crocodylians that includes one living species, the false gharial. Many more extinct species are known, extending the range of the subfamily back to the Eocene epoch. In contrast to the false gharial, which is a freshwater species that lives only in southeast Asia, extinct tomistomines had a global distribution and lived in estuaries and along coastlines. The classification of tomistomines among Crocodylia has been in flux; while traditionally thought to be within Crocodyloidea, molecular evidence indicates that they are more closely related to true gharials as members of Gavialoidea. Description Tomistomines have narrow or longirostrine snouts like gharials. The living false gharial lives in fresh water and uses its long snout and sharp teeth to catch fish, although true gharials are more adapted toward piscivory, or fish-eating. Despite the similarity with gharials, the shapes of bones in tomistomine skulls link them with crocodiles. For example, b ...
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Kentisuchus
''Kentisuchus'' is an extinct genus of gavialoid crocodylian, traditionally regarded as a member of the subfamily Tomistominae. Fossils have been found from England and France that date back to the early Eocene. The genus has also been recorded from Ukraine, but it unclear whether specimens from Ukraine are referable to ''Kentisuchus''. Species The genus ''Kentisuchus'' was erected by Charles Mook in 1955 for the species ''"Crocodylus" toliapicus'', described by Richard Owen, in 1849. William Buckland named ''"Crocodylus" spenceri'' on the basis of a partial skull found from the Isle of Sheppey in Kent, England. In 1888 Richard Lydekker considered ''"C." toliapicus'' synonymous with ''"C." champsoides'' and ''"C." arduini'', named by De Zigno, and reapplied the name ''"C." spenceri'' to all of these species. The genus name ''Kentisuchus'' was constructed only after it was realized that these specimens were clearly distinct from the genus '' Crocodylus'' and that some specime ...
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Toyotamaphimeia
''Toyotamaphimeia'' (named after Toyotama-hime) is a genus of extinct gavialid crocodylian which lived in Japan and Taiwan during the Middle Pleistocene. A specimen recovered in 1964 at Osaka University during the construction of a new science building has been dated to around 430–380 thousand years old based on the stratum in which it was found. ''Toyotamaphimeia'' was a fairly large crocodylian measuring approximately long. Two species are named, ''T. machikanensis'' from Japan and ''T. taiwanicus'' from Taiwan, both originally described as members of the genus ''Tomistoma''. History and naming The first bones belonging to ''Toyotamaphimeia'' were discovered on May 3, 1964, during the construction of a new school building on the grounds of Ôsaka University. A field survey was conducted shortly afterwards, confirming the presence of more fossils, however not yet identifying their crocodilian nature. Following the survey several digs were organized starting on 9 June 1964. Th ...
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Thecachampsa
''Thecachampsa'' is an extinct genus of gavialoid crocodylian, traditionally regarded as a member of the subfamily Tomistominae. Fossils have been found from the eastern United States in deposits of Miocene age. Those named in the 19th century were distinguished primarily by the shape of their teeth, and have since been combined with ''T. antiquus''. More recently erected species were reassigned from other genera, although their assignment to ''Thecachampsa'' has since been questioned. Description ''Thecachampsa'', like other "tomistomines" of the Oligocene and Miocene, was considerably larger than living crocodilians. Like living gharials, it had a long, slender snout. The teeth were long and recurved. Unlike its living relatives, ''Thecachampsa'' was marine, inhabiting estuaries and shallow coastal waters. Other marine fossils such as sea snail and bivalve shells, shark teeth, and barnacles have been found alongside remains of ''Thecachampsa'' and similar taxa. Species In 185 ...
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Rhamphosuchus
''Rhamphosuchus'' ("Beak crocodile") is an extinct genus of gavialid crocodylians. It lived during the Pliocene and its fossils have been found in two regions; the Siwalik Hills of Pakistan and India as well as the Sindh region of Pakistan. The genus contains two species, ''R. crassidens'' and ''R. pachyrhynchus'', with the latter originally assigned to the genus ''Gavialis''. Overview Historically, many palaeontologists estimated that it was one of the largest, if not the largest crocodylian that ever lived, reaching an estimated length of . However, a more recent study suggests that the animal may have been 8–11 m (26 to 36 ft) in length, and therefore is not the largest known crocodylian. Another crocodylian, '' Purussaurus'', from the Miocene of Peru and Brazil, is known from an equally incomplete fossil set. It is estimated to have been similar in length to the initial estimates at approximately 9.2–10.3 m (about 30–34 ft). However, this would mean that i ...
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Penghusuchus
''Penghusuchus'' is an extinct genus of gavialid crocodylian. It is known from a skeleton found in Middle to Upper Miocene rocks of Penghu Island, off Taiwan. The taxon was described in 2009 by Shan and colleagues; the type species is ''P. pani''. It may be related to two other fossil Asian gavialids: '' Toyotamaphimeia machikanensis'' of Japan and '' Hanyusuchus sinensis'' of South China. It was a medium-sized gavialid with an estimated total length of . Discovery On 25 March 2006, on the coast of Neian, Shiyu, Penghu Islands, an excavator driver Mr. Ming-Kuo Pan found a fossilized crocodylian tooth exposed in the sandstone interlayer between basaltic rocks and then dug up a whole skeleton. The skeleton is 70% complete and was found in the Yuwentao Formation of the middle Miocene (more than 10 million years ago), and its sedimentary rocks were dated as 17-15 million years ago, according to the pollen dating in the stratum, making it one of the oldest and most complete vert ...
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Paratomistoma
''Paratomistoma'' (meaning "next to or near ''Tomistoma''") is an extinct monospecific genus of gavialoid crocodylian. It is based on the holotype specimen CGM 42188, a partial posterior skull and lower jaw discovered at Wadi Hitan, Egypt, in Middle Eocene-age rocks of the Gehannam Formation. The skull is unfused but considered morphologically mature. ''Paratomistoma'' was named in 2000 by Christopher Brochu and Philip Gingerich; the type species is ''P. courti'' in honor of Nicholas Court, who found CGM 42188. They performed a phylogenetic analysis and found ''Paratomistoma'' to be a derived member of Tomistominae, related to the false gharial. It may have been a marine or coastal crocodilian. Below is a cladogram based morphological studies comparing skeletal features that shows ''Paratomistoma'' as a member of Tomistominae: Based on morphological studies of extinct taxa, the tomistomines were long thought to be classified as crocodiles and not closely re ...
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Megadontosuchus
''Megadontosuchus'' is an extinct monospecific genus of gavialoid crocodylian, traditionally regarded as a member of Tomistominae, from the middle Eocene of Italy. Fossils have been found from Monte Duello in the province of Verona. The genus is currently monotypic, with the type and only species being ''Megadontosuchus arduini''. The species was originally named in 1880, although it was assigned to the genus ''Crocodilius'' (now spelled ''Crocodylus''). The genus was first erected by paleontologist Charles C. Mook in 1955 along with the genus ''Kentisuchus'', which was also first classified as ''Crocodilius''. No holotype for ''Megadontosuchus'' was designated in 1880, and a lectotype wasn't proposed until 2007. ''Megadontosuchus'' differs from other traditional "tomistomines" in that it has a more robust rostrum, very large teeth (hence the generic name meaning ''big-toothed crocodile''), and large supratemporal fenestrae on the skull table The skull roof or the roofing bone ...
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Maroccosuchus
''Maroccosuchus zennaroi'' is an extinct gavialoid crocodylian from the Early Eocene of Morocco, traditionally regarded as a member of the subfamily Tomistominae. Below is a cladogram based on morphological studies comparing skeletal features that shows ''Maroccosuchus'' as a member of Tomistominae, related to the false gharial: Based on morphological studies of extinct taxa, the tomistomines were long thought to be classified as crocodiles and not closely related to gavialoids. However, recent molecular studies using DNA sequencing have consistently indicated that the false gharial (''Tomistoma'') (and by inference other related extinct forms in Tomistominae) actually belong to Gavialoidea (and Gavialidae). Below is a cladogram from a 2018 tip dating study by Lee & Yates simultaneously using morphological, molecular (DNA sequencing), and stratigraphic (fossil age) data that shows ''Maroccosuchus'' as an early-diverging gavialoid, more basal than the last common ancestor ...
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Gavialoidea
Gavialoidea is one of three superfamily (zoology), superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomistoma schlegelii'' are alive today, with ''Hanyusuchus'' having become extinct in the last few centuries. Extinct South American gavialoids likely Dispersal (ecology), dispersed in the mid Tertiary period, Tertiary from Africa and Asia. Fossil remains of the Puerto Rican gavialoid ''Aktiogavialis puertorisensis'' were discovered in a cave located in San Sebastián, Puerto Rico and dated to the Oligocene. This individual is thought to have crossed the Atlantic coming from Africa, indicating that this species was able to withstand saltwater. Classification Gavialoidea is Cladistics, cladistically defined as ''Gavialis gangeticus'' (the gharial) and all crocodylians closer to it than to ''Alligator mississippiensis'' (the American alligator) ...
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Gavialosuchus
''Gavialosuchus'' is an extinct genus of gavialoid crocodylian from the early Miocene of Europe. Currently only one species is recognized, as a few other species of ''Gavialosuchus'' have since been reclassified to other genera. Taxonomy The type species, ''G. eggenburgensis'', is known from the early Miocene of Austria. Two other species - ''G. americanus'' and ''G. carolinensis'' - have since been reclassified to other genera. Myrick Jr. (2001) proposed synonymizing ''Gavialosuchus americanus'' with '' Thecachampsa antiqua''. Piras ''et al.'' (2007) advocated transferring both ''G. americanus'' and ''G. carolinensis'' to ''Thecachampsa'' as distinct species of the latter genus, however. Jouve ''et al.'' (2008) retained ''G. americanus'' in ''Gavialosuchus'' and found it to be the sister group of ''G. eggenburgensis'' (''G. carolinensis'' was not discussed). However, Jouve ''et al.'' (2008) didn't test ''Thecachampsa antiqua'' in their phylogenetic analysis. Shan ''et al.'' ( ...
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Ferganosuchus
''Ferganosuchus'' is an extinct monospecific genus of gavialid crocodilian. The genus is thought to be a tomistomine, although it has been suggested that ''Ferganosuchus'' may be a more basal gavialoid. Fossils have been found in the region of Kazakhstan and Kyrgyzstan that date back the middle Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes .... The genus is well known from fairly complete specimens, unlike most other Asian tomistomines, whose remains tend to be more fragmentary.Piras, P., Delfino, M., Del Favero, L., and Kotsakis, T. (2007). Phylogenetic position of the crocodylian ''Megadontosuchus arduini'' and tomistomine palaeobiogeography. ''Acta Palaeontologica Polonica'' 52(2):315–328 References Gavialidae Eocene crocodylomorphs Eocene reptiles of Asia Pr ...
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Dollosuchoides
''Dollosuchoides'', colloquially known as the Crocodile of Maransart, is an extinct monospecific genus of gavialoid crocodilian, traditionally regarded as a member of the subfamily Tomistominae. Fossils have been found in the Brussels Formation of Maransart, Belgium and date back to the middle Eocene. The holotype, IRScNB 482, was discovered in 1915 and it was prepared during 1926–1927 by M. Hubert, J. Mehschaert and M. Jean de Kleermaeker, and also in 1927, Louis Dollo had the holotype put on display in the Museum of Natural Sciences and he intended to describe the specimen but he died in 1931 before he was able to describe it and the specimen was eventually referred to '' Dollosuchus'' by Swinton (1937)Swinton, W. E. (1937)The Crocodile of Maransart (Dollosuchus dixoni [Owen]).''Mémoire'' 80: 3–46 until it was moved to its own genus by Brochu (2007). It is currently housed in the Gand Museum in Belgium. Phylogeny Below is a cladogram based morphological stud ...
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