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Syncytia
A syncytium (; : syncytia; from Greek: σύν ''syn'' "together" and κύτος ''kytos'' "box, i.e. cell") or symplasm is a multinucleate cell that can result from multiple cell fusions of uninuclear cells (i.e., cells with a single nucleus), in contrast to a coenocyte, which can result from multiple nuclear divisions without accompanying cytokinesis. The muscle cell that makes up animal skeletal muscle is a classic example of a syncytium cell. The term may also refer to cells interconnected by specialized membranes with gap junctions, as seen in the heart muscle cells and certain smooth muscle cells, which are synchronized electrically in an action potential. The field of embryogenesis uses the word ''syncytium'' to refer to the coenocytic blastoderm embryos of invertebrates, such as ''Drosophila melanogaster''. Physiological examples Protists In protists, syncytia can be found in some rhizarians (e.g., chlorarachniophytes, plasmodiophorids, haplosporidians) and ace ...
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Multinucleate
Multinucleate cells (also known as multinucleated cells or polynuclear cells) are eukaryotic cells that have more than one nucleus, i.e., multiple nuclei share one common cytoplasm. Mitosis in multinucleate cells can occur either in a coordinated, synchronous manner where all nuclei divide simultaneously or asynchronously where individual nuclei divide independently in time and space. Certain organisms may have a multinuclear stage of their life cycle. For example, slime molds have a vegetative, multinucleate life stage called a plasmodium. Multinucleate cells, depending on the mechanism by which they are formed, can be divided into "syncytia" (formed by cell fusion) or " coenocytes" (formed by nuclear division not being followed by cytokinesis). Some bacteria, such as '' Mycoplasma pneumoniae'', a pathogen of the respiratory tract, may display multinuclear filaments as a result of a delay between genome replication and cellular division. Terminology Some biologists use ...
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Coenocyte
A coenocyte () is a multinucleate cell which can result from multiple nuclear divisions without their accompanying cytokinesis, in contrast to a syncytium, which results from cellular aggregation followed by dissolution of the cell membranes inside the mass. The word syncytium in animal embryology is used to refer to the coenocytic blastoderm of invertebrates. A coenocytic colony is referred to as a coenobium (: coenobia), and most coenobia are composed of a distinct number of cells, often as a multiple of two (4, 8, etc.). Research suggests that coenobium formation may be a defense against grazing in some species. Physiological examples Protists Diplomonads, like ''Giardia'', have two nuclei. Ciliates have cells that contain two nuclei: a macronucleus and a micronucleus. The schizont of apicomplexan parasites is a form of a coenocyte (i.e. a plasmodium in the general sense) as well as the plasmodia of microsporidian (Fungi) and myxosporidian (Metazoa) parasites. The tr ...
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Haplozoon
''Haplozoon'' (/hæploʊ’zoʊən/) are unicellular endo-parasites, primarily infecting maldanid polychaetes. They belong to Dinoflagellata but differ from typical dinoflagellates. Most dinoflagellates are free-living and possess two flagella. Instead, ''Haplozoon'' belong to a 5% minority of parasitic dinoflagellates that are not free-living. Additionally, the ''Haplozoon'' trophont stage is particularly unique due to an apparent lack of flagella. The presence of flagella or remnant structures is the subject of ongoing research. At first glance, ''Haplozoon'' also do not appear unicellular – in fact they were originally classified as a possible transitional stage between protists and multicellular organisms.Shumway, W. (1924). The genus ''Haplozoon'', Dogiel. Observations on the life history and systematic position. ''The Journal of Parasitology, 11''(2), 59-74. They have more recently been classified as compartmentalized syncytia – single cells with multiple nuclei that ...
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Tapetum (botany)
The tapetum is a specialised layer of nutritive cells found within the anther of flowering plants, located between the sporogenous tissue and the anther wall. Tapetum is important for the nutrition and development of pollen grains and a source of precursors for the pollen coat.P.L. Polowick, V.K. Sawhney 1993Differentiation of the Tapetum During Microsporogenesis in Tomato (Lycopersicon esculentum Mill.), with Special Reference to the Tapetal Cell Wall Annals of Botany, Volume 72, Issue 6, December 1993, Pages 595–605 The cells are usually bigger and normally have more than one nucleus per cell. As the sporogenous cells undergo mitosis, the nuclei of tapetal cells also divide. Sometimes, this mitosis is abnormal, which is why many cells of mature tapetum become multinucleated. Polyploidy and polyteny can also be seen sometimes. The tapetum's unusually large nuclear constitution helps it provide nutrients and regulatory molecules to the forming pollen grains. The following pro ...
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Cytokinesis
Cytokinesis () is the part of the cell division process and part of mitosis during which the cytoplasm of a single eukaryotic cell divides into two daughter cells. Cytoplasmic division begins during or after the late stages of nuclear division in mitosis and meiosis. During cytokinesis the spindle apparatus partitions and transports duplicated chromatids into the cytoplasm of the separating daughter cells. It thereby ensures that chromosome number and complement are maintained from one generation to the next and that, except in special cases, the daughter cells will be functional copies of the parent cell. After the completion of the telophase and cytokinesis, each daughter cell enters the interphase of the cell cycle. Particular functions demand various deviations from the process of symmetrical cytokinesis; for example in oogenesis in animals the ovum takes almost all the cytoplasm and organelles. This leaves very little for the resulting polar bodies, which in most spec ...
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Drosophila Melanogaster
''Drosophila melanogaster'' is a species of fly (an insect of the Order (biology), order Diptera) in the family Drosophilidae. The species is often referred to as the fruit fly or lesser fruit fly, or less commonly the "vinegar fly", "pomace fly", or "banana fly". In the wild, ''D. melanogaster'' are attracted to rotting fruit and fermenting beverages, and are often found in orchards, kitchens and pubs. Starting with Charles W. Woodworth's 1901 proposal of the use of this species as a model organism, ''D. melanogaster'' continues to be widely used for biological research in genetics, physiology, microbial pathogenesis, and Life history theory, life history evolution. ''D. melanogaster'' was the first animal to be Fruit flies in space, launched into space in 1947. As of 2017, six Nobel Prizes have been awarded to drosophilists for their work using the insect. ''Drosophila melanogaster'' is typically used in research owing to its rapid life cycle, relatively simple genetics with on ...
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Dictyostelid
The dictyostelids (Dictyostelia/Dictyostelea, ICZN, Dictyosteliomycetes, ICBN) or cellular slime molds are a group of slime molds or social amoebae. Multicellular behavior When food (normally bacteria) is readily available dictyostelids behave as individual amoebae, which feed and divide normally. However, when the food supply is exhausted, they aggregate to form a multicellular assembly, called a pseudoplasmodium, grex, or slug (not to be confused with the gastropod mollusc called a slug). The grex has a definite anterior and posterior, responds to light and temperature gradients, and has the ability to migrate. Under the correct circumstances the grex matures forming a sorocarp (fruiting body) with a stalk supporting one or more sori (balls of spores). These spores are inactive cells protected by resistant cell walls, and become new amoebae once food is available. In ''Acytostelium'', the sorocarp is supported by a stalk composed of cellulose, but in other dictyostelids ...
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Amoebozoa
Amoebozoa is a major Taxonomy (biology), taxonomic group containing about 2,400 described species of Amoeba, amoeboid protists, often possessing blunt, fingerlike, Pseudopod#Morphology, lobose pseudopods and tubular mitochondrial cristae. In traditional classification schemes, Amoebozoa is usually ranked as a phylum within either the kingdom (biology), kingdom Protista or the kingdom Protozoa. In the classification favored by the International Society of Protistologists, it is retained as an unranked "supergroup (biology), supergroup" within Eukaryota. Molecular genetics, Molecular genetic analysis supports Amoebozoa as a monophyletic clade. Modern studies of eukaryotic phylogenetic trees identify it as the sister group to Opisthokonta, another major clade which contains both fungi and animals as well as several other clades comprising some 300 species of unicellular eukaryotes. Amoebozoa and Opisthokonta are sometimes grouped together in a high-level taxon, named Amorphea. Amoeboz ...
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Acrasid
The family Acrasidae (ICZN, or Acrasiomycota, ICBN) is a family of slime molds which belongs to the excavate group Heterolobosea. The name element - comes from the Greek ''akrasia'', meaning "acting against one's judgement". This group consists of cellular slime molds. The terms "Acrasiomycota" or "Acrasiomycetes" have been used when the group was classified as a fungus ("-mycota"). In some classifications, ''Dictyostelium'' was placed in Acrasiomycetes, an artificial group of cellular slime molds, which was characterized by the aggregation of individual amoebae into a multicellular fruiting body, making it an important factor that related the acrasids to the dictyostelids. Each cell keeps its individuality even when it forms a stalk and fruiting body to reproduce. Slime molds were originally thought to be in a monophyletic group ''Mycetozoa'', with little distinction between ''Acrasis'' and ''Dictyostelids'', however scientists uncovered that they were distinct groups, and ...
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Ancient Greek
Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek Dark Ages, Dark Ages (), the Archaic Greece, Archaic or Homeric Greek, Homeric period (), and the Classical Greece, Classical period (). Ancient Greek was the language of Homer and of fifth-century Athens, fifth-century Athenian historians, playwrights, and Ancient Greek philosophy, philosophers. It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article primarily contains information about the Homeric Greek, Epic and Classical periods of the language, which are the best-attested periods and considered most typical of Ancient Greek. From the Hellenistic period (), Ancient Greek was followed by Koine Greek, which is regar ...
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Excavata
Excavata is an obsolete, extensive and diverse paraphyletic group of unicellular Eukaryota. The group was first suggested by Simpson and Patterson in 1999 and the name latinized and assigned a rank by Thomas Cavalier-Smith in 2002. It contains a variety of free-living and symbiotic protists, and includes some important parasites of humans such as ''Giardia'' and '' Trichomonas''. Excavates were formerly considered to be included in the now- obsolete Protista kingdom. They were distinguished from other lineages based on electron-microscopic information about how the cells are arranged (they have a distinctive ultrastructural identity). They are considered to be a basal flagellate lineage. On the basis of phylogenomic analyses, the group was shown to contain three widely separated eukaryote groups, the discobids, metamonads, and malawimonads. A current view of the composition of the excavates is given below, indicating that the group is paraphyletic. Except for some Eugleno ...
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