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Sporocarp (ferns)
A sporocarp is a specialised type of structure in the aquatic ferns of the order Salviniales whose primary function is the production and release of spores. Sporocarps are found only in the Salviniales, a group that is aquatic and heterosporous, but the structures are very different in the two families of the order. In the Salviniaceae family, the sporocarp is nothing more than a modified sorus, a single cluster of spore-producing tissues enclosed by a thin sphere of tissue and attached to the leaves. In the Marsileaceae (water-clover) family, the sporocarp is a more elaborate structure formed from an entire leaf whose development and form is greatly modified. These are hairy, short-stalked, bean-shaped structures (usually 3 to 8 mm in diameter) with a hardened outer covering. This outer covering is tough and resistant to drying out, allowing the spores inside to survive unfavorable conditions such as winter frost or summer desiccation. Despite this toughness, the sporocarps ...
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Pilularia Globulifera - Thomé
''Pilularia'' or pillworts is a genus of unusual ferns of family Marsileaceae distributed in North Temperate regions, Ethiopian mountains, and the southern hemisphere in Australia, New Zealand, and western South America. Depending on the taxonomic revisor, the genus contains between 3 and 6 species of small plants with thread-like leaves, and creeping rhizomes. The sporangia are borne in spherical sporocarps ("pills") which form in the axils of leaves. ''Pilularia minuta'' from SW Europe is one of the smallest of all ferns. Phylogeny Recent work from Pryer Lab accepted four species of ''Pilularia'', with a fifth, ''P. novae-zealandiae'', being conspecific with ''P. novae-hollandiae''. In addition, another species, ''P. dracomontana'', has been described from South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastl ...
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Leaf
A leaf (: leaves) is a principal appendage of the plant stem, stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, flower, and fruit collectively form the Shoot (botany), shoot system. In most leaves, the primary Photosynthesis, photosynthetic Tissue (biology), tissue is the palisade mesophyll and is located on the upper side of the blade or lamina of the leaf, but in some species, including the mature foliage of ''Eucalyptus'', palisade mesophyll is present on both sides and the leaves are said to be isobilateral. The leaf is an integral part of the stem system, and most leaves are flattened and have distinct upper (Glossary of botanical terms#adaxial, adaxial) and lower (Glossary of botanical terms#abaxial, abaxial) surfaces that differ in color, Trichome, hairiness, the number of stomata (pores that intake and output gases), the amount and ...
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Microsporangium
A microsporangium () is a sporangium that produces microspores that give rise to male gametophytes when they germinate. Microsporangia occur in all vascular plants that have heterosporic life cycles, such as seed plants, spike mosses and the aquatic fern genus ''Azolla''. In gymnosperms and angiosperm anthers, the microsporangia produce microsporocytes, the microspore mother cells, which then produce four microspores through the process of meiosis. Microsporocytes are produced in the microsporangia of gymnosperm cones and the anthers of angiosperms. They are diploid microspore mother-cells, which then produce four haploid microspores by meiosis. These become pollen grains, within which the microspores divide twice by mitosis to produce a very simple gametophyte. Heterosporous plants that produced microspores in microsporangia and megaspores in separate megasporangia evolved independently in several plant groups during the Devonian period. Fossils of these plants show that they ...
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Megaspore
Megaspores, also called macrospores, are a type of spore that is present in heterosporous plants. These plants have two spore types, megaspores and microspores. Generally speaking, the megaspore, or large spore, germinates into a female gametophyte, which produces egg cells. These are fertilized by sperm produced by the male gametophyte developing from the microspore. Heterosporous plants include seed plants (gymnosperms and flowering plants), water ferns (Salviniales), spikemosses (Selaginellaceae) and quillworts (Isoetaceae). Megasporogenesis In gymnosperms and flowering plants, the megaspore is produced inside the nucellus of the ovule. During megasporogenesis, a diploid precursor cell, the megasporocyte or megaspore mother cell, undergoes meiosis to produce initially four haploid cells (the megaspores). Angiosperms exhibit three patterns of megasporogenesis: monosporic, bisporic, and tetrasporic, also known as the ''Polygonum'' type, the ''Alisma'' type, and the ''Drusa'' ...
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Megasporangium
A sporangium (from Late Latin, ; : sporangia) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other groups form sporangia at some point in their life cycle. Sporangia can produce spores by mitosis, but in land plants and many fungi, sporangia produce genetically distinct haploid spores by meiosis. It's outdated name, sporange, is one of the few perfect rhymes for orange. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains haploid nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and germinate into haploid hyphae. Although sexual reproduction in fungi varies between phyla, for some fungi the ...
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Indusium
A sorus (: sori) is a cluster of sporangia (structures producing and containing spores) in ferns and fungi. A coenosorus (: coenosori) is a compound sorus composed of multiple, fused sori. Etymology This Neo-Latin word is from Ancient Greek σωρός (''sōrós'' 'stack, pile, heap'). Structure In lichens and other fungi, the sorus is surrounded by an external layer. In some red algae, it may take the form of depression into the thallus. In ferns, the sori form a yellowish or brownish mass on the edge or underside of a fertile frond. In some species, they are protected during development by a scale or film of tissue called the indusium (: indusia), which forms an umbrella-like cover. Life cycle significance Sori occur on the sporophyte generation, the sporangia within producing haploid meiospores. As the sporangia mature, the indusium shrivels so that spore release is unimpeded. The sporangia then burst and release the spores. As an aid to identification The shape, arrang ...
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Sporangium
A sporangium (from Late Latin, ; : sporangia) is an enclosure in which spores are formed. It can be composed of a unicellular organism, single cell or can be multicellular organism, multicellular. Virtually all plants, fungus, fungi, and many other groups form sporangia at some point in their biological life cycle, life cycle. Sporangia can produce spores by mitosis, but in land plants and many fungi, sporangia produce genetically distinct haploid spores by meiosis. It's outdated name, sporange, is one of the few perfect rhymes for Orange (colour), orange. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains Ploidy, haploid Cell nucleus, nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and g ...
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Germinate
Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an flowering plant, angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of Fungus, fungi, ferns, bacteria, and the growth of the pollen tube from the pollen grain of a seed plant. Seed plants Germination is usually the growth of a plant contained within a seed resulting in the formation of the seedling. It is also the process of reactivation of metabolic machinery of the seed resulting in the emergence of radicle and plumule. The seed of a vascular plant is a small package produced in a fruit or conifer cone, cone after the union of male and female gamete, reproductive cells. All fully developed seeds contain an embryo and, in most plant species some store of food reserves, wrapped in a seed coat. Dormant seeds are viable seeds that do not germinate because they require specific internal or environ ...
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Bean
A bean is the seed of some plants in the legume family (Fabaceae) used as a vegetable for human consumption or animal feed. The seeds are often preserved through drying (a ''pulse''), but fresh beans are also sold. Dried beans are traditionally soaked and boiled, and used in many traditional dishes throughout the world. They can be cooked in many different ways, however, including frying and baking. The unripe seedpods of some varieties are also eaten whole as green beans or '' edamame'' (immature soybean), but many fully ripened beans contain toxins like phytohemagglutinin and require cooking. Terminology The word "bean" and its Germanic cognates (e.g. German '' Bohne'') have existed in common use in West Germanic languages since before the 12th century, referring to broad beans, chickpeas, and other pod-borne seeds. This was long before the New World genus '' Phaseolus'' was known in Europe. With the Columbian exchange of domestic plants between Europe and the A ...
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Marsileaceae
Marsileaceae () is a small family of heterosporous aquatic and semi-aquatic ferns, though at first sight they do not physically resemble other ferns. The group is commonly known as the "pepperwort family" or as the "water-clover family" because the leaves of the genus ''Marsilea'' superficially resemble the leaves of a four-leaf clover. In all, the family contains three genera and 50 to 80 species with most of those belonging to ''Marsilea''. Natural history Members of the Marsileaceae are aquatic or semi-aquatic. Plants often grow in dense clumps in mud along the shores of ponds or streams, or they may grow submerged in shallow water with some of the leaves extending to float on the water surface. They grow in seasonally wet habitats, but survive the winter or dry season by losing their leaves and producing hard, desiccation-resistant reproductive structures. There are only three living genera in the family Marsileaceae. The majority of species (about 45 to 70) belon ...
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Fern
The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissues that conduct water and nutrients, and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaf, leaves called megaphylls that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled Fiddlehead fern, fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate (Polypodiidae (plant), Polypodiidae) and eusporangiate ferns, the latter group including horsetails, Psilotaceae, whisk ferns, marattioid ferns, and ophioglossoid ferns. The fern crown group, consisting of the leptosporangiates and ...
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Tissue (biology)
In biology, tissue is an assembly of similar cells and their extracellular matrix from the same embryonic origin that together carry out a specific function. Tissues occupy a Biological organisation#Levels, biological organizational level between cell (biology), cells and a complete organ (biology), organ. Accordingly, organs are formed by the functional grouping together of multiple tissues. The English word "tissue" Morphological derivation, derives from the French word "", the past participle of the verb tisser, "to weave". The study of tissues is known as histology or, in connection with disease, as histopathology. Xavier Bichat is considered as the "Father of Histology". Plant histology is Studied Space Shuttle designs, studied in both plant anatomy and Plant physiology, physiology. The classical tools for studying tissues are the Microtome#Applications, paraffin block in which tissue is embedded and then sectioned, the staining, histological stain, and the Microscope, o ...
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