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Rugose Coral
The Rugosa or rugose corals are an extinct class of solitary and colonial corals that were abundant in Middle Ordovician to Late Permian seas. Solitary rugosans (e.g., '' Caninia'', '' Lophophyllidium'', '' Neozaphrentis'', '' Streptelasma'') are often referred to as ''horn corals'' because of a unique horn-shaped chamber with a wrinkled, or rugose, wall. Some solitary rugosans reached nearly a meter (3 ft 3 in) in length. However, some species of rugose corals could form large colonies (e.g., ''Lithostrotion''). When radiating septa were present, they were usually in multiples of four, hence ''Tetracorallia'' in contrast to modern ''Hexacorallia'', colonial polyps generally with sixfold symmetry. Rugose corals have a skeleton made of calcite that is often fossilized. Like modern corals (Scleractinia), rugose corals were invariably benthic, living on the sea floor or in a reef-framework. Some symbiotic rugose corals were endobionts of Stromatoporoidea, especially in the Si ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era, and the second of twelve periods of the Phanerozoic Eon (geology), Eon. The Ordovician spans 41.6 million years from the end of the Cambrian Period Megaannum, Ma (million years ago) to the start of the Silurian Period Ma. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official per ...
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Rugose
Rugose means "wrinkled". It may refer to: * Rugosa, an extinct order of coral, whose rugose shape earned it the name * Rugose, adjectival form of rugae Species with "rugose" in their names * ''Idiosoma nigrum'', more commonly, a black rugose trapdoor spider * Rugose spiraling whitefly {{disambig ...
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Septum (coral)
In corals, a septum (plural septa) is one of the radiating vertical plates lying within the corallite A corallite is the skeletal cup, formed by an individual stony coral polyp, in which the polyp sits and into which it can retract. The cup is composed of aragonite, a crystalline form of calcium carbonate, and is secreted by the polyp. Corallit ... wall. Outside the corallite wall these plates are known as costae (singular costa). The septa may be thick, thin or vary in size. They may have teeth which range from needle-like to blade-like and are often characteristic of different genera. References {{reflist Cnidarian anatomy ...
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Corallite
A corallite is the skeletal cup, formed by an individual stony coral polyp, in which the polyp sits and into which it can retract. The cup is composed of aragonite, a crystalline form of calcium carbonate, and is secreted by the polyp. Corallites vary in size, but in most colonial corals they are less than in diameter. The inner surface of the corallite is known as the calyx. The vertical blades inside the calyx are known as septa SEPTA, the Southeastern Pennsylvania Transportation Authority, is a regional public transportation authority that operates bus, rapid transit, commuter rail, light rail, and electric trolleybus services for nearly four million people througho ... and in some species, these ridges continue outside the corallite wall as costae. Where there is no corallite wall, the blades are known as septocostae. The septa, costae and septocostae may have ornamentation in the form of teeth and may be thick, thin or variable in size. Sometimes there are palifor ...
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Palaeozoic
The Paleozoic ( , , ; or Palaeozoic) Era is the first of three geological eras of the Phanerozoic Eon. Beginning 538.8 million years ago (Ma), it succeeds the Neoproterozoic (the last era of the Proterozoic Eon) and ends 251.9 Ma at the start of the Mesozoic Era. The Paleozoic is subdivided into six geologic periods (from oldest to youngest), Cambrian, Ordovician, Silurian, Devonian, Carboniferous and Permian. Some geological timescales divide the Paleozoic informally into early and late sub-eras: the Early Paleozoic consisting of the Cambrian, Ordovician and Silurian; the Late Paleozoic consisting of the Devonian, Carboniferous and Permian. The name ''Paleozoic'' was first used by Adam Sedgwick (1785–1873) in 1838 to describe the Cambrian and Ordovician periods. It was redefined by John Phillips (1800–1874) in 1840 to cover the Cambrian to Permian periods. It is derived from the Greek ''palaiós'' (παλαιός, "old") and ''zōḗ'' (ζωή, "life") meaning "ancient l ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) ...
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Stromatoporoidea
Stromatoporoidea is an extinct clade of sea sponges common in the fossil record from the Middle Ordovician to the Late Devonian.Stock, C.W. 2001, Stromatoporoidea, 1926–2000: ''Journal of Paleontology'', v. 75, p. 1079–1089. They can be characterized by their densely layered calcite skeletons lacking Sponge spicule, spicules. Stromatoporoids were among the most abundant and important reef-builders of their time, living close together in flat biostromes or elevated bioherms on soft tropical carbonate platforms. Externally, some species have raised bumps (mamelons) and star-shaped crevices (astrorhizae), which together help vent exhalant water away from the living surface. Internally, stromatoporoids have a mesh-like skeletal system combining extensive horizontal layers (laminae), vertical rods (pillars), and boxy spaces (galleries), along with other features. The most common growth forms range from laminar (flattened) to domical (dome-shaped). Spheroidal, finger-like, or tree-l ...
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Endobiont
An endosymbiont or endobiont is an organism that lives within the body or cells of another organism. Typically the two organisms are in a mutualistic relationship. Examples are nitrogen-fixing bacteria (called rhizobia), which live in the root nodules of legumes, single-cell algae inside reef-building corals, and bacterial endosymbionts that provide essential nutrients to insects. Endosymbiosis played key roles in the development of eukaryotes and plants. Roughly 2.2 billion years ago an archaeon absorbed a bacterium through phagocytosis, that eventually became the mitochondria that provide energy to almost all living eukaryotic cells. Approximately 1 billion years ago, some of those cells absorbed cyanobacteria that eventually became chloroplasts, organelles that produce energy from sunlight. Approximately 100 million years ago, a lineage of amoeba in the genus ''Paulinella'' independently engulfed a cyanobacterium that evolved to be functionally synonymous with traditional ch ...
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Benthic
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". Organisms living in this zone are called benthos and include microorganisms (e.g., bacteria and fungi) as well as larger invertebrates, such as crustaceans and polychaetes. Organisms here, known as bottom dwellers, generally live in close relationship with the substrate and many are permanently attached to the bottom. The benthic boundary layer, which includes the bottom layer of water and the uppermost layer of sediment directly influenced by the overlying water, is an integral part of the benthic zone, as it greatly influences the biological activity that takes place there. Examples of contact soil layers include sand bottoms, rocky outcrops, coral, and bay mud. Description Oceans The benthic region of the ocean begins at the ...
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Scleractinia
Scleractinia, also called stony corals or hard corals, are marine animals in the phylum Cnidaria that build themselves a hard skeleton. The individual animals are known as polyps and have a cylindrical body crowned by an oral disc in which a mouth is fringed with tentacles. Although some species are solitary, most are colonial. The founding polyp settles and starts to secrete calcium carbonate to protect its soft body. Solitary corals can be as much as across but in colonial species the polyps are usually only a few millimetres in diameter. These polyps reproduce asexually by budding, but remain attached to each other, forming a multi-polyp colony of clones with a common skeleton, which may be up to several metres in diameter or height according to species. The shape and appearance of each coral colony depends not only on the species, but also on its location, depth, the amount of water movement and other factors. Many shallow-water corals contain symbiont unicellular organis ...
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