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Polykrikoid
The Polykrikaceae (also known as Polykrikidae) are a family of athecate dinoflagellates of the order Gymnodiniales. Members of the family are known as polykrikoids. The family contains two genera: ''Polykrikos'' and '' Pheopolykrikos''. Characteristics The most distinctive feature of polykrikoids is their formation of multinucleate "pseudocolonies" consisting of an even number of subunit zooids. The two genera differ in number of nuclei; possessing two nuclei regardless of the number of zooids is a synapomorphy for ''Polykrikos'', whereas ''Pheopolykrikos'' possess equal numbers of nuclei and zooids. Along with the Warnowiaceae (warnowiids), polykrikoids are known for possessing unusually complex subcellular structures. In particular, an extrusome complex of two organelles called the nematocyst and taeniocyst is considered a synapomorphy for ''Polykrikos''. Molecular phylogenetics studies suggest some inconsistency in the taxonomy of this group, particularly in the assignmen ...
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Polykrikos
''Polykrikos'' (from Greek “poly” - many, and “krikos” – ring or circle) is one of the genera of family Polykrikaceae that includes athecate pseudocolony-forming dinoflagellates. ''Polykrikos'' are characterized by a sophisticated ballistic apparatus, named the nematocyst-taeniocyst complex, which allows species to prey on a variety of organisms. ''Polykrikos'' have been found to regulate algal blooms as they feed on toxic dinoflagellates. However, there is also some data available on Polykrikos being toxic to fish. History of knowledge ''Polykrikos'' was first seen in 1868 by Uljanin and was mistakenly considered as a metazoan larva of a turbellarian flatworms. In 1873 Butschili re-examined the specimen and concluded that the cell was an unusual ciliate, and Bergh later, in 1881, clarified ''Polykrikos'' dinoflagellate affinities. Morphology ''Polykrikos'' is a colony of zooids (units of a colonial organism) that carry out simultaneous functions of a whole cell. ...
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Polykrikos Kofoidii
''Polykrikos kofoidii'' is a species of phagotrophic marine pseudocolonial dinoflagellates that can capture and engulf other protist prey, including the toxic dinoflagellate, '' Alexandrium tamarense''. '' P. kofoidii'' is of scientific interest due to its status as a predator of other dinoflagellates, a behavior that is significant in the control of algal bloom An algal bloom or algae bloom is a rapid increase or accumulation in the population of algae in fresh water or marine water systems. It is often recognized by the discoloration in the water from the algae's pigments. The term ''algae'' encompass ...s. It has a complex life cycle of both vegetative (asexual) and sexual reproduction complicated by its pseudocolonial structure. References Protists described in 1914 Gymnodiniales Dinoflagellate species {{dinoflagellate-stub ...
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Nematocyst (dinoflagellate)
A nematocyst is a subcellular structure or organelle containing extrusive filaments found in two families of athecate dinoflagellates (a group of unicellular eukaryotes), the Warnowiaceae and Polykrikaceae. It is distinct from the similar subcellular structures found in the cnidocyte cells of cnidarians, a group of multicellular organisms including jellyfish and corals; such structures are also often called nematocysts (alternatively, ''cnidocysts'' or ''cnidae''), and cnidocytes are sometimes referred to as nematocytes. It is unclear whether the relationship between dinoflagellate and cnidarian nematocysts is a case of convergent evolution or common descent, although molecular evidence has been interpreted as supporting an endosymbiotic origin for cnidarian nematocysts. In polykrikoids the nematocyst is found associated with another extrusive organelle called the taeniocyst, a complex that has been described as synapomorphic for the genus '' Polykrikos''. The full range of f ...
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Warnowiaceae
The Warnowiaceae are a family of athecate dinoflagellates (a diverse group of unicellular eukaryotes). Members of the family are known as warnowiids. The family is best known for a light-sensitive subcellular structure known as the ocelloid, a highly complex arrangement of organelles with a structure directly analogous to the eyes of multicellular organisms. The ocelloid has been shown to be composed of multiple types of endosymbionts, namely mitochondria and at least one type of plastid. Habitat and life cycle Warnowiids are found in marine plankton but are very rare in most plankton samples. Little is known about their life histories because they cannot be cultured in the laboratory, and samples obtained from the natural environment do not survive well under laboratory conditions. Studies of wild samples have found evidence of distinctive structures called trichocysts in warnowiid cell vacuoles, suggesting that their prey might be other dinoflagellates. Despite the complexit ...
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Plankton
Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a crucial source of food to many small and large aquatic organisms, such as bivalves, fish, and baleen whales. Marine plankton include bacteria, archaea, algae, protozoa, microscopic fungi, and drifting or floating animals that inhabit the saltwater of oceans and the brackish waters of estuaries. fresh water, Freshwater plankton are similar to marine plankton, but are found in lakes and rivers. Mostly, plankton just drift where currents take them, though some, like jellyfish, swim slowly but not fast enough to generally overcome the influence of currents. Although plankton are usually thought of as inhabiting water, there are also airborne versions that live part of their lives drifting in the at ...
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Organelle
In cell biology, an organelle is a specialized subunit, usually within a cell (biology), cell, that has a specific function. The name ''organelle'' comes from the idea that these structures are parts of cells, as Organ (anatomy), organs are to the Human body, body, hence ''organelle,'' the suffix ''-elle'' being a diminutive. Organelles are either separately enclosed within their own lipid bilayers (also called membrane-bounded organelles) or are spatially distinct functional units without a surrounding lipid bilayer (non-membrane bounded organelles). Although most organelles are functional units within cells, some functional units that extend outside of cells are often termed organelles, such as cilia, the flagellum and archaellum, and the trichocyst (these could be referred to as membrane bound in the sense that they are attached to (or bound to) the membrane). Organelles are identified by microscopy, and can also be purified by cell fractionation. There are many types of organe ...
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Molecular Phylogenetics
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetics, phylogenetic analysis is expressed in a phylogenetic tree. Molecular phylogenetics is one aspect of molecular systematics, a broader term that also includes the use of molecular data in Taxonomy (biology), taxonomy and biogeography. Molecular phylogenetics and molecular evolution correlate. Molecular evolution is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tre ...
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Taxonomy (biology)
In biology, taxonomy () is the science, scientific study of naming, defining (Circumscription (taxonomy), circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxon, taxa (singular: taxon), and these groups are given a taxonomic rank; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain (biology), domain, kingdom (biology), kingdom, phylum (''division'' is sometimes used in botany in place of ''phylum''), class (biology), class, order (biology), order, family (biology), family, genus, and species. The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, having developed a ranked system known as Linnaean taxonomy for categorizing organisms. With advances in the theory, data and analytical technology of biological systematics, the Linnaean system has transfo ...
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Light Microscopy
Microscopy is the technical field of using microscopes to view subjects too small to be seen with the naked eye (objects that are not within the resolution range of the normal eye). There are three well-known branches of microscopy: optical, electron, and scanning probe microscopy, along with the emerging field of X-ray microscopy. Optical microscopy and electron microscopy involve the diffraction, reflection, or refraction of electromagnetic radiation/electron beams interacting with the specimen, and the collection of the scattered radiation or another signal in order to create an image. This process may be carried out by wide-field irradiation of the sample (for example standard light microscopy and transmission electron microscopy) or by scanning a fine beam over the sample (for example confocal laser scanning microscopy and scanning electron microscopy). Scanning probe microscopy involves the interaction of a scanning probe with the surface of the object of interest. The de ...
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Benthic
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". Organisms living in this zone are called benthos and include microorganisms (e.g., bacteria and fungi) as well as larger invertebrates, such as crustaceans and polychaetes. Organisms here, known as bottom dwellers, generally live in close relationship with the substrate and many are permanently attached to the bottom. The benthic boundary layer, which includes the bottom layer of water and the uppermost layer of sediment directly influenced by the overlying water, is an integral part of the benthic zone, as it greatly influences the biological activity that takes place there. Examples of contact soil layers include sand bottoms, rocky outcrops, coral, and bay mud. Description Oceans The benthic region of the ocean begins at the ...
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Photosynthetic
Photosynthesis ( ) is a Biological system, system of biological processes by which Photoautotrophism, photosynthetic organisms, such as most plants, algae, and cyanobacteria, convert light energy, typically from sunlight, into the chemical energy necessary to fuel their metabolism. ''Photosynthesis'' usually refers to oxygenic photosynthesis, a process that produces oxygen. Photosynthetic organisms store the chemical energy so produced within intracellular organic compounds (compounds containing carbon) like sugars, glycogen, cellulose and starches. To use this stored chemical energy, an organism's cells metabolize the organic compounds through cellular respiration. Photosynthesis plays a critical role in producing and maintaining the atmospheric oxygen, oxygen content of the Earth's atmosphere, and it supplies most of the biological energy necessary for complex life on Earth. Some bacteria also perform anoxygenic photosynthesis, which uses bacteriochlorophyll to split hydrogen ...
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