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Polycotylidae
Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widespread during the early Late Cretaceous. Several species survived into the final stage of the Cretaceous, the Maastrichtian. With their short necks and large elongated heads, they resemble the pliosaurs, but closer phylogenetic studies indicate that they share many common features with the Leptocleididae and Elasmosauridae. They have been found worldwide, with specimens reported from New Zealand, Australia, Japan, Morocco, the US, Canada, Eastern Europe, and South America. Phylogeny Cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... after Albright, Gillette and ...
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Mauriciosaurus
''Mauriciosaurus'' (meaning "Mauricio ernández Garza'sreptile") is a genus of polycotylid plesiosaur from the Late Cretaceous of Mexico. It contains a single species, ''M. fernandezi'' (also named after Fernández Garza), described in 2017 by Eberhard Frey and colleagues from a single well-preserved juvenile specimen about long. Morphologically, it is overall most similar to the polycotyline polycotylids ''Trinacromerum'' and ''Dolichorhynchops''. However, several features separate ''Mauriciosaurus'' from all other polycotylids, warranting the naming of a new genus. These include the sophisticated pattern of ridges on the bottom of the parasphenoid bone on its palate; the narrow openings in the palate bordered by the pterygoid bones; the lack of perforations in the surface of the coracoid; and the highly unusual arrangement of gastralia, or belly ribs, which is only otherwise seen in the non-polycotylid ''Cryptoclidus''. Partially as a consequence of the oxygen-poor environ ...
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Sulcusuchus
''Sulcusuchus'' is a genus of polycotylid plesiosaur from the Late Campanian of Argentina. Description The type species ''Sulcusuchus erraini'' was named in 1990 by Zulma Brandoni de Gasparini and Luis Spalletti. It was at first considered to have been a dyrosaurid crocodile, hence its generic name: "trough crocodile". Its holotype, MPEF 650, was at Jacobacci, Río Negro Province, uncovered in a layer of the late Campanian La Colonia Formation. It consists of a posterior left mandible fragment of about half a meter long with an associated jaw joint condyle of the skull, lower braincase, rear palate and middle snout fragment. The outer side of the mandible shows the trough, which would be exceptional for a crocodile. The mandibula and pterygoids show grooves that may have housed electro-sensitive organs.O’Gorman, J.P. & Gasparini, Z., 2013, "Revision of ''Sulcusuchus erraini'' (Sauropterygia, Polycotylidae) from the Upper Cretaceous of Patagonia, Argentina", ''Alcheringa'' ...
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Trinacromerum
''Trinacromerum'' is an extinct genus of sauropterygian reptile, a member of the polycotylid plesiosaurs. It contains two species, ''T. bentonianum'' and ''T. kirki''. Specimens have been discovered in the Late Cretaceous fossil deposits of what is now modern Kansas and Manitoba. Description ''Trinacromerum'' was long. Its teeth show that it fed on small fish. The long flippers of ''Trinacromerum'' enabled it to achieve high swimming speeds. Its physical appearance was described by Richard Ellis as akin to a "four-flippered penguin."Ellis, 190 Its name means "three tipped femur". Classification Below is a cladogram of polycotylid relationships from Ketchum & Benson, 2011. See also * List of plesiosaur genera * Timeline of plesiosaur research This timeline of plesiosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, taxonomic revisions, and cultural portrayals of plesiosaurs, an order of marine reptil ...
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Dolichorhynchops Bonneri
''Dolichorhynchops'' is an extinct genus of polycotylid plesiosaur from the Late Cretaceous (early Turonian to late Campanian stage) of North America, containing three species, ''D. osborni'', ''D. bonneri'' and ''D. tropicensis'', as well as a questionably referred fourth species, ''D. herschelensis''. ''Dolichorhynchops'' was an oceangoing prehistoric reptile. Its Greek generic name means "long-nosed face". Discovery and species ''D. osborni'' The holotype specimen of ''Dolichorhynchops osborni'' was discovered in the upper Smoky Hill Chalk Logan County, Kansas, by George F. Sternberg, as a teenager, in around 1900. The remains were collected by him and his father, Charles H. Sternberg, and then sold to the University of Kansas (Lawrence, Kansas). KUVP 1300 was prepared and mounted by H.T. Martin under the supervision of Dr. Samuel Wendell Williston, who described and named it in 1902. A more detailed description and photographs were provided by ). The specimen has been on d ...
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Dolichorhynchops
''Dolichorhynchops'' is an extinct genus of polycotylid plesiosaur from the Late Cretaceous (early Turonian to late Campanian stage) of North America, containing three species, ''D. osborni'', ''D. bonneri'' and ''D. tropicensis'', as well as a questionably referred fourth species, ''D. herschelensis''. ''Dolichorhynchops'' was an oceangoing prehistoric reptile. Its Greek generic name means "long-nosed face". Discovery and species ''D. osborni'' The holotype specimen of ''Dolichorhynchops osborni'' was discovered in the upper Smoky Hill Chalk Logan County, Kansas, by George F. Sternberg, as a teenager, in around 1900. The remains were collected by him and his father, Charles H. Sternberg, and then sold to the University of Kansas (Lawrence, Kansas). KUVP 1300 was prepared and mounted by H.T. Martin under the supervision of Dr. Samuel Wendell Williston, who described and named it in 1902. A more detailed description and photographs were provided by ). The specimen has been on ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', " chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Ear ...
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Leptocleididae
Leptocleididae is a family of small-sized plesiosaurs that lived during the Early Cretaceous period (early Berriasian to early Albian stage). They had small bodies with small heads and short necks. '' Leptocleidus'' and '' Umoonasaurus'' had round bodies and triangle-shaped heads. Leptocleidids have been found in what were shallow nearshore, freshwater and brackish habitats. Hilary F. Ketchum and Roger B. J. Benson (2010), transferred ''Brancasaurus ''Brancasaurus'' (meaning "Branca's lizard") is a genus of plesiosaur which lived in a freshwater lake in the Early Cretaceous of what is now North Rhine-Westphalia, Germany. With a long neck possessing vertebrae bearing distinctively-shaped "s ...'', '' Kaiwhekea'', '' Nichollssaura'' and '' Thililua'' to this family. However, Ketchum and Benson (2011) reassigned ''Kaiwhekea'' and ''Thililua'' to their original positions, as an elasmosaurid and a polycotylid, respectively. Phylogeny Cladogram based on Ketchum and Benson (2011 ...
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Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma (million years ago). The Albian is preceded by the Aptian and followed by the Cenomanian. Stratigraphic definitions The Albian Stage was first proposed in 1842 by Alcide d'Orbigny. It was named after Alba, the Latin name for River Aube in France. A Global Boundary Stratotype Section and Point (GSSP), ratified by the IUGS in 2016, defines the base of the Albian as the first occurrence of the planktonic foraminiferan '' Microhedbergella renilaevis'' at the Col de Pré-Guittard section, Arnayon, Drôme, France. The top of the Albian Stage (the base of the Cenomanian Stage and Upper Cretaceous Series) is defined as the place where the foram species '' Rotalipora globotruncanoides'' first appears in the stratigraphic column. Th ...
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Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from . The Maastrichtian was preceded by the Campanian and succeeded by the Danian (part of the Paleogene and Paleocene). The Cretaceous–Paleogene extinction event (formerly known as the Cretaceous–Tertiary extinction event) occurred at the end of this age. In this mass extinction, many commonly recognized groups such as non-avian dinosaurs, plesiosaurs and mosasaurs, as well as many other lesser-known groups, died out. The cause of the extinction is most commonly linked to an asteroid about wide colliding with Earth, ending the Cretaceous. Stratigraphic definitions Definition The Maastrichtian was introduced into scientific literature by Belgian geologist André Hubert Dumont in 1849, after studying rock strata of the ...
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Early Cretaceous
The Early Cretaceous (geochronological name) or the Lower Cretaceous ( chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java- Manihiki- Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kergue ...
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Pliosaur
Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toothed jaws, commonly known as pliosaurs. More primitive non-thalassophonean pliosauroids resembled pleisiosaurs in possessing relatively long necks and smaller heads. They originally included only members of the family Pliosauridae, of the order Plesiosauria, but several other genera and families are now also included, the number and details of which vary according to the classification used. The distinguishing characteristics are a short neck and an elongated head, with larger hind flippers compared to the fore flippers, the opposite of the plesiosaurs. They were carnivorous and their long and powerful jaws carried many sharp, conical teeth. Pliosaurs range from 4 to 15 metres and more in length. Their prey may have included fish, shar ...
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Elasmosauridae
Elasmosauridae is an extinct family of plesiosaurs, often called elasmosaurs. They had the longest necks of the plesiosaurs and existed from the Hauterivian to the Maastrichtian stages of the Cretaceous, and represented one of the two groups of plesiosaurs present at the end of the Cretaceous alongside Polycotylidae. Their diet mainly consisted of crustaceans and molluscs. Description The earliest elasmosaurids were mid-sized, about . In the Late Cretaceous, elasmosaurids grew as large as , such as ''Styxosaurus'', '' Albertonectes'', and ''Thalassomedon''. Their necks were the longest of all the plesiosaurs, with anywhere between 32 and 76 (''Albertonectes'') cervical vertebrae. They weighed up to several tons. Classification Early three-family classification Though Cope had originally recognized ''Elasmosaurus'' as a plesiosaur, in an 1869 paper he placed it, with '' Cimoliasaurus'' and '' Crymocetus'', in a new order of sauropterygian reptiles. He named the group S ...
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