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Osteostracan
The class Osteostraci (meaning "bony shells") is an extinct taxon of bony-armored jawless fish, termed "ostracoderms", that lived in what is now North America, Europe and Russia from the Middle Silurian to Late Devonian. Anatomically speaking, the osteostracans, especially the Devonian species, were among the most advanced of all known agnathans. This is due to the development of paired fins, and their complicated cranial anatomy. The osteostracans were more similar to lampreys than to jawed vertebrates in possessing two pairs of semicircular canals in the inner ear, as opposed to the three pairs found in the inner ears of jawed vertebrates. They are thought to be the sister-group to pituriaspids, and together, these two taxa of jawless vertebrates are the sister-group of gnathostomes. Several synapomorphies support this hypothesis, such as the presence of: sclerotic ossicles, paired pectoral fins, a dermal skeleton with three layers (a basal layer of isopedin, a middle layer o ...
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Osteostraci Janvier
The class (biology), class Osteostraci (meaning "bony shells") is an extinct taxon of bony-armored jawless fish, termed "ostracoderms", that lived in what is now North America, Europe and Russia from the Wenlock epoch, Middle Silurian to Late Devonian. Anatomically speaking, the osteostracans, especially the Devonian species, were among the most advanced of all known agnathans. This is due to the development of paired fins, and their complicated cranial anatomy. The osteostracans were more similar to lampreys than to jawed vertebrates in possessing two pairs of semicircular canals in the inner ear, as opposed to the three pairs found in the inner ears of jawed vertebrates. They are thought to be the sister-group to Pituriaspida, pituriaspids, and together, these two taxa of jawless vertebrates are the sister-group of Gnathostomata, gnathostomes. Several synapomorphies support this hypothesis, such as the presence of: sclerotic ossicles, paired pectoral fins, a dermal skeleton with ...
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Cephalaspis
''Cephalaspis'' (from , 'head' and , 'shield') is a possibly monotypic genus of extinct osteostracan agnathan vertebrate. It was a trout-sized detritivorous fish that lived in the early Devonian. Description Like its relatives, ''Cephalaspis'' was heavily armored, presumably to defend against predatory placoderms and eurypterids, as well as to serve as a source of calcium for metabolic functions in calcium-poor freshwater environments. It had sensory patches along the rim and center of its head shield, which were used to sense for worms and other burrowing organisms in the mud. Diet Because its mouth was situated directly beneath its head, ''Cephalaspis'' was thought of as being a bottom-feeder, akin to a heavily armoured catfish or sturgeon. It moved its plow-like head from side to side, ''Cephalaspis'' easily stirring sand and dust into the water, along with revealing the hiding places of its prey, digging up worms or crustaceans hidden in the mud and algae, as well as sif ...
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Wladysagitta
''Wladysagitta'' is an extinct genus of osteostracan jawless fish that existed during the lower Devonian period of what is now Podolia, Ukraine. This taxon was named in honor of Polish paleontologist Dr. Władysław Zych (1899–1981), and from the Latin sagitta, meaning arrow, which is in reference to the arrow-like shape of its skull.Voichyshyn, Victor. (2006). New osteostracans from the Lower Devonian terrigenous deposits of Podolia, Ukraine. Acta Palaeontologica Polonica. 51. Description ''Wladysagitta'', like other osteostracans, possesses very advanced features for an agnathan; namely, an armored headshield with complicated cranial anatomy, paired fins and rows of plated scales. It is remarkable among osteostracans for possessing a rostral process, a beaklike structure unusual for this class, which usually possesses blunted, horseshoe-shaped headshields. It is also distinct for being ornamented by a number of unusually prominent, thornlike tubercles on its headshield as pre ...
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Zenaspida
Zenaspidida is an extinct order of osteostracans, a group of jawless stem- gnathostomes. They possessed a distinct headshield, which varied in width to length ratio by species. Description The head shield is dome-shaped and extremely large in comparison to the main body. The abdominal section of this shield has a less developed median dorsal crest. As a rule for this order, the nasohypophysial opening is larger than the nasal division. The pineal plate seen in other osteostracans is barely developed or completely absent. The median dorsal field is notably broad, and the lateral fields are widened in the posterior, but reach back no further than the proximal section of the dorsal surface of the cornual processes. The ornamentation on the head shield can have singular, large tubercles, or groups of tubercles which range in size. This is often used to speciate. Classification The cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "characte ...
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Agnatha
Agnatha (; ) or jawless fish is a paraphyletic infraphylum of animals in the subphylum Vertebrata of the phylum Chordata, characterized by the lack of jaws. The group consists of both extant taxon, living (Cyclostomi, cyclostomes such as hagfishes and lampreys) and Extinction, extinct clades (e.g. conodonts and Cephalaspidomorphi, cephalaspidomorphs, among others). They are sister taxon, sister to vertebrates with jaws known as gnathostomes, who evolution, evolved from jawless ancestors during the early Silurian by developing folding joint, articulations in the first pairs of gill arches. Sequencing, Molecular data, both from rRNA and from mtDNA as well as Embryology, embryological data, strongly supports the hypothesis that both groups of living agnathans, hagfishes and lampreys, are more closely related to each other than to Gnathostomata, jawed fish, forming the Class (biology), superclass Cyclostomi. The oldest fossil agnathans appeared in the Cambrian. Living jawless fish c ...
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Benneviaspidida
Benneviaspidida is an order of osteostracan jawless fishes which lived in the Early Devonian. The fishes in this order have a flat headshield and are dorsoventrally depressed. The first canal to lateral sensory field bifurcates near the orbit. Phylogeny The cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ... below is adapted from Sansom (2009): References External links Benneviaspidida at PalaeosOsteostraci by Philippe JanvierEarly Devonian osteostracans from Russia Osteostraci Prehistoric jawless fish orders Devonian jawless fish Early Devonian first appearances Early Devonian taxonomic orders Early Devonian extinctions {{Devonian-jawless-fish-stub ...
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Thyestiida
Thyestiida is an order of bony-armored jawless fish in the extinct vertebrate class Osteostraci The class Osteostraci (meaning "bony shells") is an extinct taxon of bony-armored jawless fish, termed " ostracoderms", that lived in what is now North America, Europe and Russia from the Middle Silurian to Late Devonian. Anatomically speaking, ....The origin and early evolution of the Osteostraci (Vertebrata): A phylogeny for the Thyestiida. Robert S. Sansom, Journal of Systematic Palaeontology, Volume 6, Issue 3, 2008, pages 317-332, References External links Thyestiidaat fossilworks.org (retrieved 21 April 2016) Osteostraci Prehistoric jawless fish orders {{Paleo-jawless-fish-stub ...
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Ostracoderm
Ostracodermi () or ostracoderms is an informal group of vertebrate animals that include all armored jawless fish of the Paleozoic Era. The term does not often appear in classifications today because it is paraphyletic (excluding jawed fishes and possibly the cyclostomes if anaspids are closer to them) and thus does not correspond to one evolutionary lineage. However, the term is still used as an informal way of loosely grouping together the armored jawless fishes. An innovation of ostracoderms was the use of gills not for feeding, but exclusively for respiration. Earlier chordates with gill precursors used them for both respiration and feeding. Ostracoderms had separate pharyngeal gill pouches along the side of the head, which were permanently open with no protective operculum. Unlike invertebrates that use ciliated motion to move food, ostracoderms used their muscular pharynx to create a suction that pulled small and slow-moving prey into their mouths. Swiss anatomist L ...
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Gnathostomata
Gnathostomata (; from Ancient Greek: (') 'jaw' + (') 'mouth') are jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all extant vertebrates, including all living bony fishes (both ray-finned and lobe-finned, including their terrestrial tetrapod relatives) and cartilaginous fishes, as well as extinct prehistoric fish such as placoderms and acanthodians. Most gnathostomes have retained ancestral traits like true teeth, a stomach, and paired appendages ( pectoral and pelvic fins, limbs, wings, etc.). Other traits are elastin, horizontal semicircular canal of the inner ear, myelinated neurons, and an adaptive immune system which has discrete secondary lymphoid organs (spleen and thymus) and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene. It is now assumed that Gnathostomata evolved from ancestors that already possessed two pairs ...
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Ateleaspididae
Ateleaspididae is a prehistoric jawless fish family in the class Osteostraci The class Osteostraci (meaning "bony shells") is an extinct taxon of bony-armored jawless fish, termed " ostracoderms", that lived in what is now North America, Europe and Russia from the Middle Silurian to Late Devonian. Anatomically speaking, ....Sansom, R. S. (2009). "Phylogeny, classification and character polarity of the Osteostraci (Vertebrata)". Journal of Systematic Palaeontology 7: 95–11. References External links Osteostraci Prehistoric jawless fish families {{Paleo-jawless-fish-stub ...
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