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Neoophora
Neoophora is a group of rhabditophoran flatworms with ectolecithal eggs, i.e., yolk is not present in the egg as in most animals, but rather is secreted by accessory glands called vitellaria or yolk glands. These glands have the same embryonic origin as the ovaries, but usually constitute a separate organ in adult animals. The monophyly of Neoophora is disputed, since some recent molecular studies indicated that the most basal order, Lecithoepitheliata, is more closely related to Polycladida, a non-neoophoran order, than to other neoophorans. This would imply that the ectolecithal condition would have evolved twice. The clade formed by all other neoophorans, excluding Lecithoepitheliata, is usually called Euneoophora Trepaxonemata (from ''trepa''-, spiral + axoneme) is a subclass of the Platyhelminthes or flatworms.Ehlers, U. (1985). Das Phylogenetische System der Plathelminthes. Stuttgart: G. Fischer. It includes all parasitic flatworms (clade Neodermata) .... References ...
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Lecithoepitheliata
The Lecithoepitheliata are an order of rhabditophoran flatworms. They are free-living worms, found in freshwater, soil, and marine environments. However, it is still poorly known their roles in the natural food web. Description Members of the order Lecithoepitheliata are distinguished from other flatworms by the presence of four nerve cords and the fact that the ovary forms a single structure that produces both the eggs (ovocytes) and nourishing yolk cells (vitellocytes). The vitellocytes form a follicle around the ovocyte, hence the name of the group, which means "with a yolk epithelium". In most other rhabditophorans, yolk cells, where present at all, are typically formed in glands derived from the ovaries, but separate from them, called vitellaria. Other diagnostic features of this order include the presence of a sharp stylet on the end of the penis, and a simple, unbranched, intestine. The penis lacks a proper pore and is protruded through the mouth. Phylogeny The order Lec ...
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Rhabdocoela
Rhabdocoela is an order of flatworms in the class Rhabditophora with about 1700 species described worldwide. The order was first described in 1831 by Christian Gottfried Ehrenberg. Most of rhabdocoels are free-living organisms, but some live symbiotically with other animals. Description Although Rhabdocoela is a highly supported group in molecular studies, there is no clear morphological synapomorphy that unites them. All rhabdocoels have a bulbous pharynx, but this is shared with other flatworm groups, such as Neodermata, Lecithoepitheliata and some species of Prolecithophora. Some possibly identified synapomorphies are found in the ultrastructure of the protonephridial system, but similar constructions exist in other groups. Another possible apomorphy is found in the ultrastructure of the sperm, which has a dense heel on the basal bodies during spermiogenesis, but some groups have lost this feature. Classification Rhabdocoels were traditionally classified in two group ...
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Proseriata
Proseriata is an order of free-living flatworms in the class Rhabditophora with over 400 species described worldwide. Description Proseriate flatworms are minute aquatic organisms, usually with an elongate body. There are no strong synapomorphies for the group, but it is supported by molecular studies. The suggested synapomorphies include the lack of lamellated rhabdites, an otherwise synapomorphy of Rhabditophora, and some features of the ultrastructure of the protonephridia and the cilia of epidermal cells. Ecology Most proseriates occur in marine environments, at the coastal zones, and are particularly common in high-energy habitats with medium to coarse sediments, sometimes being the dominant animal group in such areas. They are also common in brackish water, but few species occur in freshwater. Most proseriates are carnivores or scavengers, but some species are ectoparasitic on marine crustaceans. As they occur in very high densities in some habitats, they may have a co ...
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Prolecithophora
The ''Prolecithophora'' are an order consisting of an estimated 300 species of small (typically 0.2 – 12 mm, one species up to 50 mm), active, aquatic flatworms. The order lacks a common English name. Most species are shaped like an elongated, stylized droplet, and are opaque white or yellow; they frequently have contrasting bands or spots in colors, such as purple, yellow, red, or brown. They have no to three (normally two) pairs of pigment-cup eyes, and well-developed tactile and chemoreceptor senses. With few exceptions, species are protandric hermaphrodites with internal fertilization. Egg capsules are, according to species, glued to various hard surfaces; the young hatch as miniature copies of their parents. Ecology All prolecithophorans are aquatic, with most living in the oceans. Some species, especially those living in freshwater, are predators and scavengers, but many marine species are associated with colonial animals such as bryozoans or live as symbiont ...
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Fecampiida
Fecampiida is an order of flatworms in the class Rhabditophora. It is a considerably recent clade, erected after molecular studies. Description The order Fecampiida, as currently defined, was erected based on molecular studies. They all are parasitic organisms and are united by a similar development of the basal bodies during spermiogenesis. Classification Three families of Fecampiida were initially classified in different flatworm orders: Urastomatidae and Genostomatidae were considered prolecithophorans, while Fecampiidae was considered a rhabdocoel. When the genus '' Notentera'' was discovered, its relationship with Fecampiidae was clear based on morphology, and both groups were united under Fecampiida. Further ultrastructural studies suggested that Urastomatidae and Genostomatidae were closely related to Fecampiidae and Notenteridae, which was confirmed by molecular studies. Due to similarities in the protonephridial flame bulb, sperm and spermiogenesis, as well a ...
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Tricladida
A planarian is one of the many flatworms of the traditional class Turbellaria. It usually describes free-living flatworms of the order Tricladida (triclads), although this common name is also used for a wide number of free-living platyhelminthes. Planaria are common to many parts of the world, living in both saltwater and freshwater ponds and rivers. Some species are terrestrial and are found under logs, in or on the soil, and on plants in humid areas. The triclads are characterized by triply branched intestine and anteriorly situated ovaries, next to the brain. Today the order Tricladida is split into three suborders, according to their phylogenetic relationships: Maricola, Cavernicola and Continenticola. Formerly, the Tricladida was split according to habitats: Maricola, which is marine; Paludicola which inhabits freshwater; and Terricola, which is land-dwelling. Planaria exhibit an extraordinary ability to regenerate lost body parts. For example, a planarian split len ...
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Bothrioplanida
''Bothrioplana'' is a genus of freshwater flatworms, the sole genus in the family Bothrioplanidae and order Bothrioplanida. Description Species of ''Bothrioplana'' are small organisms, usually measuring 2–3 mm in length, but reaching up to 7 mm. The body is transparent, elongate, with a round posterior end and a truncate front end, and lacks eyes. The mouth lies on the ventral side at about the middle of the body, as in most turbellarians, and has a short pharynx. The intestine is tripartite, similarly to what occurs in triclads, having one anterior and two posterior branches. The posterior branches unite behind the copulatory apparatus and form a single branch. The reproductive system has a pair of testes and a pair of ovaries, as well as a series of vitellaria (yolk-producing glands) that extend along the entire body. Many specimens have the male reproductive system highly reduced or even absent. As a result, parthenogenesis may occur. Ecology Individuals of ''Bothriop ...
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Neodermata
Neodermata is a clade of rhabditophoran flatworms containing the parasitic groups Trematoda, Monogenea and Cestoda. Description All neodermatans are parasites, in many groups having a free-swimming larval stage. The most striking feature uniting all neodermatans is that the ciliated epidermis (typical of most flatworms) is cast off in adult worms, being replaced by a syncytium called tegument or ''neodermis''. Other characters found in all neodermatans are related to the anatomy of the protonephridium and the rootlets of epidermal locomotory cilia. Phylogeny Currently, the monophyly of Neodermata is undisputed, being supported by both morphological and molecular data. It is clear that they evolved from free-living flatworms (turbellarians), but their sister-group was for a long time a matter of debate. The first attempts to reconstruct the phylogeny of flatworms, based on morphological evidence, considered Rhabdocoela Rhabdocoela is an order of flatworms in the class Rhab ...
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Rhabditophora
Rhabditophora (from ''rhabdito''-, rhabdite + Greek -φορος ''phoros'' bearer, i.e., "rhabdite bearers") is a class of flatworms. It includes all parasitic flatworms (clade Neodermata) and most free-living species that were previously grouped in the now obsolete class Turbellaria. Therefore, it contains the majority of the species in the phylum Platyhelminthes, excluding only the catenulids, to which they appear to be the sister group. The clade Rhabditophora was originally erected by Ulrich Ehlers in 1985Ehlers, U. (1985) ''Phylogenetic relationships within the Platyhelminthes''. ''In'' S. Conway Morris; J. D. George; R. Gibson; H. M. Platt (Eds.), ''The origins and relationships of lower invertebrates''. Oxford, Clarendon Press, p. 143-158. based on morphological analyses and its monophyly was later confirmed by molecular studies. Description Rhabditophorans are characterized by the presence of lamellated rhabdites, rodlike granules secreted in the cells of the epi ...
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Flatworms
The flatworms, flat worms, Platyhelminthes, or platyhelminths (from the Greek πλατύ, ''platy'', meaning "flat" and ἕλμινς (root: ἑλμινθ-), ''helminth-'', meaning "worm") are a phylum of relatively simple bilaterian, unsegmented, soft-bodied invertebrates. Unlike other bilaterians, they are acoelomates (having no body cavity), and have no specialized circulatory and respiratory organs, which restricts them to having flattened shapes that allow oxygen and nutrients to pass through their bodies by diffusion. The digestive cavity has only one opening for both ingestion (intake of nutrients) and egestion (removal of undigested wastes); as a result, the food cannot be processed continuously. In traditional medicinal texts, Platyhelminthes are divided into Turbellaria, which are mostly non-parasitic animals such as planarians, and three entirely parasitic groups: Cestoda, Trematoda and Monogenea; however, since the turbellarians have since been proven not to be ...
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Yolk
Among animals which produce eggs, the yolk (; also known as the vitellus) is the nutrient-bearing portion of the egg whose primary function is to supply food for the development of the embryo. Some types of egg contain no yolk, for example because they are laid in situations where the food supply is sufficient (such as in the body of the host of a parasitoid) or because the embryo develops in the parent's body, which supplies the food, usually through a placenta. Reproductive systems in which the mother's body supplies the embryo directly are said to be matrotrophic; those in which the embryo is supplied by yolk are said to be lecithotrophic. In many species, such as all birds, and most reptiles and insects, the yolk takes the form of a special storage organ constructed in the reproductive tract of the mother. In many other animals, especially very small species such as some fish and invertebrates, the yolk material is not in a special organ, but inside the egg cell. ...
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