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Monoicous
Monoicy () is a sexual system in haploid plants (mainly bryophytes) where both sperm and eggs are produced on the same gametophyte, in contrast with dioicy, where each gametophyte produces only sperm or eggs but never both.Crandall-Stotler, B.J. & Bartholomew-Began, S.E. (2007). ''Morphology of Mosses (Phylum Bryophyta)''. In: Flora of North America Editorial Committee, eds. (1993+). ''Flora of North America North of Mexico''. 16+ vols. New York and Oxford. Volume 27, 2007.Bell, P.R. & Helmsley, A.R. (2000). ''Green plants, their origin and diversity'' (2nd ed.). Cambridge University Press. Both monoicous () and dioicous gametophytes produce gametes in gametangia by mitosis rather than meiosis, so that sperm and eggs are genetically identical with their parent gametophyte. It has been suggested that monoicy may have benefits in dry habitats where the ability to produce sporophytes is limited due to lack of water. Monoicy is similar to, and often conflated with, monoecy, w ...
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Sexual System
A sexual system is a pattern of sex allocation or a distribution of male and female function across organisms in a species. Terms like reproductive system and mating system have also been used as synonyms. The distinction between sexual systems is not always clear due to phenotypic plasticity. Sexual systems are viewed as a key factor for genetic variation and reproductive success, and may have also led to the origin or extinction of certain species. Interests in sexual systems go back to Darwin, who found that barnacles contain species that are androdioecious and some that are dioecious. Types of sexual systems In plants there are monomorphic sexual systems where a species has hermaphrodite, male and/or female flowers on the same plant. Monomorphic sexual systems include monoecy, gynomonoecy, andromonoecy and trimonoecy. There are also dimorphic sexual systems like dioecy, gynodioecy and androdioecy. Mixed sexual systems are where hermaphrodites coexist with single se ...
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Bryophyte
The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants ( embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in size and prefer moist habitats although they can survive in drier environments. The bryophytes consist of about 20,000 plant species. Bryophytes produce enclosed reproductive structures (gametangia and sporangia), but they do not produce flowers or seeds. They reproduce sexually by spores and asexually by fragmentation or the production of gemmae. Though bryophytes were considered a paraphyletic group in recent years, almost all of the most recent phylogenetic evidence supports the monophyly of this group, as originally classified by Wilhelm Schimper in 1879. The term ''bryophyte'' comes . Terminology The term "Bryophyta" was first suggested by Braun in 1864. G.M. Smith placed this group between Algae and Pteridophyta. Features ...
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Dioicy
Dioicy () is a sexual system where archegonia and antheridia are produced on separate gametophytes. It is one of the two main sexual systems in bryophytes. Both dioicous () and monoicous gametophytes produce gametes in gametangia by mitosis rather than meiosis, so that sperm and eggs are genetically identical with their parent gametophyte. Description Dioicy promotes outcrossing. Sexual dimorphism is commonly found in dioicous species. However, according to Bernard Goffinet sexual dimorphism is rare in dioicous moss species. Dioicy is correlated with reduced sporophyte production, due to spatial separation of male and female colonies, scarcity or absence of males. The term dioecy is meaningless for bryophytes because it refers to the sexuality of sporophytes. Nonetheless dioecy and dioicy are comparable in many respects. Etymology The words dioicous and di(o)ecious are derived from οἶκος or οἰκία and δι- (di-), twice, double. (''(o)e'' is the Latin ...
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Bryophytes
The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in size and prefer moist habitats although they can survive in drier environments. The bryophytes consist of about 20,000 plant species. Bryophytes produce enclosed reproductive structures (gametangia and sporangia), but they do not produce flowers or seeds. They reproduce sexually by spores and asexually by fragmentation or the production of gemmae. Though bryophytes were considered a paraphyletic group in recent years, almost all of the most recent phylogenetic evidence supports the monophyly of this group, as originally classified by Wilhelm Schimper in 1879. The term ''bryophyte'' comes . Terminology The term "Bryophyta" was first suggested by Braun in 1864. G.M. Smith placed this group between Algae and Pteridophyta. Features Th ...
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Dioicous
Dioicy () is a sexual system where archegonia and antheridia are produced on separate gametophytes. It is one of the two main sexual systems in bryophytes. Both dioicous () and monoicous gametophytes produce gametes in gametangia by mitosis rather than meiosis, so that sperm and eggs are genetically identical with their parent gametophyte. Description Dioicy promotes outcrossing. Sexual dimorphism is commonly found in dioicous species. However, according to Bernard Goffinet sexual dimorphism is rare in dioicous moss species. Dioicy is correlated with reduced sporophyte production, due to spatial separation of male and female colonies, scarcity or absence of males. The term dioecy is meaningless for bryophytes because it refers to the sexuality of sporophytes. Nonetheless dioecy and dioicy are comparable in many respects. Etymology The words dioicous and di(o)ecious are derived from οἶκος or οἰκία and δι- (di-), twice, double. (''(o)e'' is the Latin way ...
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Genes
In biology, the word gene (from , ; "... Wilhelm Johannsen coined the word gene to describe the Mendelian units of heredity..." meaning ''generation'' or ''birth'' or ''gender'') can have several different meanings. The Mendelian gene is a basic unit of heredity and the molecular gene is a sequence of nucleotides in DNA that is transcribed to produce a functional RNA. There are two types of molecular genes: protein-coding genes and noncoding genes. During gene expression, the DNA is first copied into RNA. The RNA can be directly functional or be the intermediate template for a protein that performs a function. The transmission of genes to an organism's offspring is the basis of the inheritance of phenotypic traits. These genes make up different DNA sequences called genotypes. Genotypes along with environmental and developmental factors determine what the phenotypes will be. Most biological traits are under the influence of polygenes (many different genes) as well as g ...
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Antheridia
An antheridium is a haploid structure or organ producing and containing male gametes (called ''antherozoids'' or sperm). The plural form is antheridia, and a structure containing one or more antheridia is called an androecium. Androecium is also the collective term for the stamens of flowering plants. Antheridia are present in the gametophyte phase of cryptogams like bryophytes and ferns. Many algae and some fungi, for example ascomycetes and water moulds, also have antheridia during their reproductive stages. In gymnosperms and angiosperms, the male gametophytes have been reduced to pollen grains and in most of these the antheridia have been reduced to a single generative cell within the pollen grain. During pollination, this generative cell divides and gives rise to sperm cells. The female counterpart to the antheridium in cryptogams is the archegonium, and in flowering plants is the gynoecium. An antheridium typically consists of sterile cells and spermatogenous tissue. The ...
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Archegonia
An archegonium (pl: archegonia), from the ancient Greek ''ἀρχή'' ("beginning") and ''γόνος'' ("offspring"), is a multicellular structure or organ of the gametophyte phase of certain plants, producing and containing the ovum or female gamete. The corresponding male organ is called the antheridium. The archegonium has a long neck canal or venter and a swollen base. Archegonia are typically located on the surface of the plant thallus, although in the hornworts they are embedded. Bryophytes In bryophytes and other cryptogams sperm reach the archegonium by swimming in water films, whereas in Pinophyta and Angiosperms the pollen are delivered by wind or animal vectors and the sperm are delivered by means of a pollen tube. In the moss '' Physcomitrella patens'', archegonia are not embedded but are located on top of the leafy gametophore (s. Figure). The Polycomb protein FIE is expressed in the unfertilized egg cell (right) as the blue colour after GUS staining reveals ...
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Inbreeding
Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genetic disorders and other consequences that may arise from expression of deleterious or recessive traits resulting from incestuous sexual relationships and consanguinity. Animals avoid incest only rarely. Inbreeding results in homozygosity, which can increase the chances of offspring being affected by recessive traits. In extreme cases, this usually leads to at least temporarily decreased biological fitness of a population (called inbreeding depression), which is its ability to survive and reproduce. An individual who inherits such deleterious traits is colloquially referred to as ''inbred''. The avoidance of expression of such deleterious recessive alleles caused by inbreeding, via inbreeding avoidance mechanisms, is the main selective reason for ou ...
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Radula (plant)
''Radula'' is a genus of liverwort, and is the only genus in family Radulaceae. It is a leafy liverwort. The appearance of the plants are as a scaly, green surface on the trunk of a tree, log or rock in a sheltered, moist out-door environment. The leaves are rounded, overlapping and consist of two unequal lobes. The smaller lobe is folded beneath the larger one. The oldest fossil species is ''Radula cretacea'' from the Cenomanian aged Burmese amber of Myanmar, belonging to the subgenus '' Odontoradula.'' Molecular evidence suggests that the genus arose during the Triassic, around 227.8 Ma, and the crown group began to diversify during the Early Jurassic, around 176.3 Ma. Species include: * '' Radula boninensis'' * '' Radula carringtonii'' J.B.Jack * '' Radula cavifolia'' * '' Radula complanata'' * '' Radula deflexilobula'' Promma, L.N.Zhang et R.L.Zhu * '' Radula demissa'' M.A.M.Renner, 2013 * '' Radula javanica'' * '' Radula jonesii'' Bouman, Dirkse & Yamada * '' Radula kojana ...
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Zygosity
Zygosity (the noun, zygote, is from the Greek "yoked," from "yoke") () is the degree to which both copies of a chromosome or gene have the same genetic sequence. In other words, it is the degree of similarity of the alleles in an organism. Most eukaryotes have two matching sets of chromosomes; that is, they are diploid. Diploid organisms have the same loci on each of their two sets of homologous chromosomes except that the sequences at these loci may differ between the two chromosomes in a matching pair and that a few chromosomes may be mismatched as part of a chromosomal sex-determination system. If both alleles of a diploid organism are the same, the organism is homozygous at that locus. If they are different, the organism is heterozygous at that locus. If one allele is missing, it is hemizygous, and, if both alleles are missing, it is nullizygous. The DNA sequence of a gene often varies from one individual to another. These gene variants are called alleles. While some g ...
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Conocephalum Conicum
''Conocephalum conicum'', also known as the great scented liverwort or snakeskin liverwort, is a liverwort species in the genus '' Conocephalum''. ''C. conicum'' is part of the ''Conocephalum conicum'' complex, which includes several cryptic species. The name ''C. conicum'' refers to the cone-shaped archegoniaphore, which bear sporangia. Habitat and distribution ''C. conicum'' is one of the most common liverworts in northern hemisphere and is widely distributed throughout Canada. ''C. conicum'' is found in open woodlands, sandy banks, wet rocks or cliffs and moist soils and is strongly associated with calcareous substrates. Morphology Thalli ''C. conicum'' is the largest of the thalloid liverworts, growing up to 20 cm long. The thalli can grow to 17 mm wide. The thalli are very strong-smelling, with purplish margins; a dark green, leathery surface; flat and smooth. There is a set of lines running along the thalli's surface. The air pores, which are found betwee ...
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