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Megaspore
Megaspores, also called macrospores, are a type of spore that is present in heterosporous plants. These plants have two spore types, megaspores and microspores. Generally speaking, the megaspore, or large spore, germinates into a female gametophyte, which produces egg cells. These are fertilized by sperm produced by the male gametophyte developing from the microspore. Heterosporous plants include seed plants (gymnosperms and flowering plants), water ferns (Salviniales), spikemosses ( Selaginellaceae) and quillworts ( Isoetaceae). Megasporogenesis In gymnosperms and flowering plants, the megaspore is produced inside the nucellus of the ovule. During megasporogenesis, a diploid precursor cell, the megasporocyte or megaspore mother cell, undergoes meiosis to produce initially four haploid cells (the megaspores). Angiosperms exhibit three patterns of megasporogenesis: monosporic, bisporic, and tetrasporic, also known as the ''Polygonum'' type, the ''Alisma'' type, and the ''Drusa ...
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Ovule
In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the '' integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the female gametophyte (formed from a haploid megaspore) in its center. The female gametophyte — specifically termed a ''megagametophyte''— is also called the ''embryo sac'' in angiosperms. The megagametophyte produces an egg cell for the purpose of fertilization. The ovule is a small structure present in the ovary. It is attached to the placenta by a stalk called a funicle. The funicle provides nourishment to the ovule. Location within the plant In flowering plants, the ovule is located inside the portion of the flower called the gynoecium. The ovary of the gynoecium produces one or more ovules and ultimately becomes the fruit wall. Ovules are attached to the placenta in the ovary through a stalk-like structure known as a ''funiculus'' ...
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Heterosporous
Heterospory is the production of spores of two different sizes and sexes by the sporophytes of land plants. The smaller of these, the microspore, is male and the larger megaspore is female. Heterospory evolved during the Devonian period from isospory independently in several plant groups: the clubmosses, the ferns including the arborescent horsetails, and progymnosperms. This occurred as part of the process of evolution of the timing of sex differentiation.Sussex, I.M. (1966) The origin and development of heterospory in vascular plants. Chapter 9 in ''Trends in Plant morphogenesis'', ed. by E.G. Cutter, Longmans. Origin of heterospory Heterospory evolved due to natural selection that favoured an increase in propagule size compared with the smaller spores of homosporous plants. Heterosporous plants, similar to anisosporic plants, produce two different sized spores in separate sporangia that develop into separate male and female gametophytes. It is proposed that the em ...
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Gametophyte
A gametophyte () is one of the two alternation of generations, alternating multicellular organism, multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the Sexual reproduction of plants, sexual phase in the life cycle of plants and algae. It develops sex organs that produce gametes, haploid sex cells that participate in fertilization to form a diploid zygote which has a double set of chromosomes. Cell division of the zygote results in a new diploid multicellular organism, the second stage in the life cycle known as the sporophyte. The sporophyte can produce haploid spores by meiosis that on germination produce a new generation of gametophytes. Algae In some multicellular green algae (''Ulva lactuca'' is one example), red algae and brown algae, sporophytes and gametophytes may be externally indistinguishable (isomorphic). In ''Ulva (genus), Ulv ...
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Glossary Of Botanical Terms
This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary of leaf morphology. For other related terms, see Glossary of phytopathology, Glossary of lichen terms, and List of Latin and Greek words commonly used in systematic names. A B ...
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Spore
In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. Under favourable conditions the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote which develops into ...
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Double Fertilization
Double fertilization is a complex fertilization mechanism of flowering plants (angiosperms). This process involves the joining of a female gametophyte ( megagametophyte, also called the embryo sac) with two male gametes (sperm). It begins when a pollen grain adheres to the stigma of the carpel, the female reproductive structure of a flower. The pollen grain then takes in moisture and begins to germinate, forming a pollen tube that extends down toward the ovary through the style. The tip of the pollen tube then enters the ovary and penetrates through the micropyle opening in the ovule. The pollen tube proceeds to release the two sperm in the megagametophyte. The cells of an unfertilized ovule are 8 in number and arranged in the form of 3+2+3 (from top to bottom) i.e. 3 antipodal cells, 2 polar central cells, 2 synergids & 1 egg cell. One sperm fertilizes the egg cell and the other sperm combines with the two polar nuclei of the large central cell of the megagametophyte. The ha ...
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Microspore
Microspores are land plant spores that develop into male gametophytes, whereas megaspores develop into female gametophytes. The male gametophyte gives rise to sperm cells, which are used for fertilization of an egg cell to form a zygote. Megaspores are structures that are part of the alternation of generations in many seedless vascular cryptogams, all gymnosperms and all angiosperms. Plants with heterosporous life cycles using microspores and megaspores arose independently in several plant groups during the Devonian period. Microspores are haploid, and are produced from diploid microsporocytes by meiosis. Morphology The microspore has three different types of wall layers. The outer layer is called the perispore, the next is the exospore, and the inner layer is the endospore. The perispore is the thickest of the three layers while the exospore and endospore are relatively equal in width. Seedless vascular plants In heterosporous seedless vascular plants, modified leaves ca ...
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Gymnosperm
The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, '' Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνόσπερμος ( el, γυμνός, translit=gymnos, lit=naked, label=none and el, σπέρμα, translit=sperma, lit=seed, label=none), literally meaning 'naked seeds'. The name is based on the unenclosed condition of their seeds (called ovules in their unfertilized state). The non-encased condition of their seeds contrasts with the seeds and ovules of flowering plants (angiosperms), which are enclosed within an ovary. Gymnosperm seeds develop either on the surface of scales or leaves, which are often modified to form cones, or solitary as in yew, ''Torreya'', '' Ginkgo''. Gymnosperm lifecycles involve alternation of generations. They have a dominant diploid sporophyte phase and a reduced haploid gametophyte phase which is dependen ...
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Megasporangium
A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cycle. Sporangia can produce spores by mitosis, but in nearly all land plants and many fungi, sporangia are the site of meiosis and produce genetically distinct haploid spores. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains haploid nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and germinate into haploid hyphae. Although sexual reproduction in fungi varies between phyla, for some fungi the sporangium plays an indirect role in sexual reprodu ...
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Salviniales
The order Salviniales (formerly known as the Hydropteridales and including the former Marsileales) is an order of ferns in the class Polypodiopsida. Description Salviniales are all aquatic and differ from all other ferns in being heterosporous, meaning that they produce two different types of spores (megaspores and microspores) that develop into two different types of gametophytes (female and male gametophytes, respectively), and in that their gametophytes are endosporic, meaning that they never grow outside the spore wall and cannot become larger than the spores that produced them. The megasporangia each produce a single megaspore. In being heterosporus with endosporic gametophytes they are more similar to seed plants than to other ferns. The fertile and sterile leaves are dimorphic, taking on a different shape, and leaves bear anastomosing veins. Aerenchyma is frequently present in roots, shoots, and petioles (leaf stalks). The ferns of this order vary radically in form ...
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