Malawimonad
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Malawimonad
Malawimonads (order Malawimonadida) are a small group of microorganisms with a basal position in the evolutionary tree of eukaryotes, containing only three recognized species. They're considered part of a paraphyletic group known as "Excavata". Evolution It is clear that the malawimonads are a monophyletic clade at the base of Eukaryota, but there is no consensus on the specific relationships between other basal groups, such as Discoba, Metamonada, Ancyromonadida and Podiata. The sister group to Malawimonadida varies greatly between analyses. Some phylogenetic analyses find Malawimonadida as the sister group to Podiata. Other analyses recover Malawimonadida as the sister group of Discoba or Metamonada. Very few modern analyses recover the three clades, Malawimonadida, Discoba and Metamonada, as a monophyletic Excavata. Taxonomy History The malawimonads were first described as order Malawimonadida in 2003 by Thomas Cavalier-Smith. In 2013 they were also described as a class ...
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Imasidae
''Imasa'' is a genus of marine flagellates containing the single species ''Imasa heleensis'', discovered in 2020 in the Solomon Islands. It is the first marine member of a basal group of eukaryotes known as Malawimonadida. It is the only member of the family Imasidae. Etymology The genus name ''Imasa'' is the name of the research vessel used to obtain the cells of ''Imasa heleensis'' that were isolated. It refers to the pause taken by a crocodile between its emergence from the water and its attack. This pause between actions is resembled by the movement of the anterior flagellum of ''Imasa'', which often appears to pause near the point between the extension and recovery strokes. The specific epithet ''heleensis'' refers to ‘Hele’, which is the name of the isles containing the collection site of ''Imasa heleensis'' at Solomon Islands. As with the genus name, it is common among the indigenous languages of the region. Characteristics Morphology ''Imasa heleensis'' was isolat ...
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Excavata
Excavata is an obsolete, extensive and diverse paraphyletic group of unicellular Eukaryota. The group was first suggested by Simpson and Patterson in 1999 and the name latinized and assigned a rank by Thomas Cavalier-Smith in 2002. It contains a variety of free-living and symbiotic protists, and includes some important parasites of humans such as ''Giardia'' and '' Trichomonas''. Excavates were formerly considered to be included in the now- obsolete Protista kingdom. They were distinguished from other lineages based on electron-microscopic information about how the cells are arranged (they have a distinctive ultrastructural identity). They are considered to be a basal flagellate lineage. On the basis of phylogenomic analyses, the group was shown to contain three widely separated eukaryote groups, the discobids, metamonads, and malawimonads. A current view of the composition of the excavates is given below, indicating that the group is paraphyletic. Except for some Eugleno ...
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Discoba
Excavata is an obsolete, extensive and diverse Paraphyly, paraphyletic group of unicellular Eukaryote, Eukaryota. The group was first suggested by Simpson and Patterson in 1999 and the name latinized and assigned a rank by Thomas Cavalier-Smith in 2002. It contains a variety of free-living and symbiotic protists, and includes some important parasites of humans such as ''Giardia'' and ''Trichomonas''. Excavates were formerly considered to be included in the now- obsolete Protist, Protista kingdom. They were distinguished from other lineages based on electron-microscopic information about how the cells are arranged (they have a distinctive ultrastructural identity). They are considered to be a Basal_(phylogenetics), basal flagellate lineage. On the basis of phylogenomic analyses, the group was shown to contain three widely separated eukaryote groups, the discobids, metamonads, and malawimonads. A current view of the composition of the excavates is given below, indicating that th ...
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Malawimonadida
Malawimonads (order Malawimonadida) are a small group of microorganisms with a basal position in the evolutionary tree of eukaryotes, containing only three recognized species. They're considered part of a paraphyletic group known as "Excavata". Evolution It is clear that the malawimonads are a monophyletic clade at the base of Eukaryota, but there is no consensus on the specific relationships between other basal groups, such as Discoba, Metamonada, Ancyromonadida and Podiata. The sister group to Malawimonadida varies greatly between analyses. Some phylogenetic analyses find Malawimonadida as the sister group to Podiata. Other analyses recover Malawimonadida as the sister group of Discoba or Metamonada. Very few modern analyses recover the three clades, Malawimonadida, Discoba and Metamonada, as a monophyletic Excavata. Taxonomy History The malawimonads were first described as order Malawimonadida in 2003 by Thomas Cavalier-Smith. In 2013 they were also described as a cl ...
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Malawimonadidae
Malawimonadidae is a family of unicellular eukaryotes The eukaryotes ( ) constitute the domain of Eukaryota or Eukarya, organisms whose cells have a membrane-bound nucleus. All animals, plants, fungi, seaweeds, and many unicellular organisms are eukaryotes. They constitute a major group of ... of outsize importance in understanding eukaryote phylogeny. Malawimonadidae is a unique and relatively lesser-known group of flagellate protists belonging to the phylum Malawimonada. These organisms are single-celled eukaryotes that are found in various aquatic habitats, including freshwater environments and marine systems. Several features make Malawimonadidae unique: Evolutionary position: Malawimonads are considered to be one of the earliest branching lineages of eukaryotes. This makes them particularly interesting for researchers studying the early evolution of eukaryotic cells and their organelles. Feeding mechanism: Malawimonads are known to be bacterivorous, meaning th ...
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Gefionella Okellyi
''Gefionella'' is a genus of excavate protists belonging to the family Malawimonadidae, a basal group in the evolution of eukaryotes. It is a monotypic genus, with only the species ''Gefionella okellyi'', described in 2018. The genus is named after the Norse goddess Gefjon, while the species is named after the scientist Charles J. O'Kelly, a pioneer in the ultrastructural and phylogenetic investigation of excavate flagellates A flagellate is a cell or organism with one or more whip-like appendages called flagellum, flagella. The word ''flagellate'' also describes a particular construction (or level of organization) characteristic of many prokaryotes and eukaryote .... References Taxa described in 2018 Protist genera {{Excavata-stub ...
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Tsukubea
''Tsukubamonas'' is a unicellular heterotrophic, biflagellated excavate of the Discoba clade (along with jakobids, euglenozoans and percolozoans) with only one species known, ''Tsukubamonas globosa''. It inhabits fresh-water, feeds on bacteria, and can exist as a vegetative cell or cyst. The cells are characterised with a spherical or semi-spherical shape, are highly vacuolated with thin subsurface vesicles and the absence of a contractile vacuole, tubular cristae in its mitochondria, and two flagella of an apparatus with five main structures (four basal bodies, three major microtubule roots, four major fibres, one microtubule organization center, and several internal microtubules). ''Tsukubamonas'' is notable for having a backwards right root, a differentiation of its anterior root orientation, and for having a lack of a left root. Other notable differences are in the morphology of its singlet root and associated fibre, the lack of flagella vanes, and in its cytoskeletal stru ...
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Plants
Plants are the eukaryotes that form the kingdom Plantae; they are predominantly photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with cyanobacteria to produce sugars from carbon dioxide and water, using the green pigment chlorophyll. Exceptions are parasitic plants that have lost the genes for chlorophyll and photosynthesis, and obtain their energy from other plants or fungi. Most plants are multicellular, except for some green algae. Historically, as in Aristotle's biology, the plant kingdom encompassed all living things that were not animals, and included algae and fungi. Definitions have narrowed since then; current definitions exclude fungi and some of the algae. By the definition used in this article, plants form the clade Viridiplantae (green plants), which consists of the green algae and the embryophytes or land plants ( hornworts, liverworts, mosses, lycophytes, ferns, conifers and other ...
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Chromalveolates
Chromalveolata was a eukaryote supergroup present in a major classification of 2005, then regarded as one of the six major groups within the eukaryotes. It was a refinement of the kingdom Chromista, first proposed by Thomas Cavalier-Smith in 1981. Chromalveolata was proposed to represent the organisms descended from a single secondary endosymbiosis involving a red alga and a bikont. The plastids in these organisms are those that contain chlorophyll c. However, the monophyly of the Chromalveolata has been rejected. Thus, two papers published in 2008 have phylogenetic trees in which the chromalveolates are split up, and recent studies continue to support this view. Groups and classification Historically, many chromalveolates were considered plants, because of their cell walls, photosynthetic ability, and in some cases their morphological resemblance to the land plants (Embryophyta). However, when the five-kingdom system (proposed in 1969) took prevalence over the animal–p ...
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Jakobea
Jakobida is an order (sole order in the class Jakobea) of free-living, heterotrophic, flagellar eukaryotes in the clade Discoba. They are small (less than 15  μm), and can be found in aerobic and anaerobic environments. The order Jakobida, believed to be monophyletic, consists of only twenty species at present, and was classified as a group in 1993. There is ongoing research into the mitochondrial genomes of jakobids, which are unusually large and bacteria-like, evidence that jakobids may be important to the evolutionary history of eukaryotes. Molecular phylogenetic evidence suggests strongly that jakobids are most closely related to Heterolobosea and Euglenozoa. Description Jakobids have two flagella, inserted in the anterior end of the cell, and, like other members of order Excavata, have a ventral feeding groove and associated cytoskeleton support. The posterior flagella has a dorsal vane and is aligned within the ventral groove, where it generates a current that the ...
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Light Microscopy
Microscopy is the technical field of using microscopes to view subjects too small to be seen with the naked eye (objects that are not within the resolution range of the normal eye). There are three well-known branches of microscopy: optical, electron, and scanning probe microscopy, along with the emerging field of X-ray microscopy. Optical microscopy and electron microscopy involve the diffraction, reflection, or refraction of electromagnetic radiation/electron beams interacting with the specimen, and the collection of the scattered radiation or another signal in order to create an image. This process may be carried out by wide-field irradiation of the sample (for example standard light microscopy and transmission electron microscopy) or by scanning a fine beam over the sample (for example confocal laser scanning microscopy and scanning electron microscopy). Scanning probe microscopy involves the interaction of a scanning probe with the surface of the object of interest. The de ...
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Euglenozoa
Euglenozoa are a large group of flagellate Discoba. They include a variety of common free-living species, as well as a few important parasites, some of which infect humans. Euglenozoa are represented by four major groups, ''i.e.,'' Kinetoplastea, Diplonemea, Euglenida, and Symbiontida. Euglenozoa are unicellular, mostly around in size, although some euglenids get up to long. Structure Most euglenozoa have two flagella, which are inserted parallel to one another in an apical or subapical pocket. In some these are associated with a cytostome or mouth, used to ingest bacteria or other small organisms. This is supported by one of three sets of microtubules that arise from the flagellar bases; the other two support the dorsal and ventral surfaces of the cell. Some other euglenozoa feed through absorption, and many euglenids possess chloroplasts, the only eukaryotes outside Diaphoretickes to do so without performing kleptoplasty, and so obtain energy through photosynthesis. The ...
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